Euphorbia oblongata |
Euphorbia carunculata |
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Balkan spurge, egg-leaf spurge, oblong spurge |
sand-dune sandmat or spurge |
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Habit | Herbs, perennial, with woody taproot. | Herbs, annual, with taproot. |
Stems | erect, unbranched or densely branching, 80 cm, often densely villous (especially young stems and pleiochasial branches). |
prostrate, spreading and lanky or occasionally mat-forming, ± succulent, 70–150 cm, glabrous. |
Leaves | petiole absent; blade oblong to narrowly obovate or lanceolate, 15–70 × 6–25 mm, base rounded or truncate, margins finely serrulate, apex obtuse, mucronulate, surfaces glabrous; venation inconspicuously pinnate, midvein prominent. |
opposite; stipules usually distinct, occasionally connate basally (primarily at distal nodes), usually divided into 2–5 subulate to subulate-filiform segments, occasionally forming narrow deltate segments (primarily at distal nodes), 0.8–1.8 mm, glabrous; petiole 3.1–6.3 mm, glabrous; blade ovate to elliptic-oblong, 5–26 × 4–12 mm, base subsymmetric to symmetric, rounded to cuneate, margins entire, apex usually mucronate, rarely acute or obtuse, surfaces glabrous; pinnately veined. |
Involucre | cupulate to slightly turbinate, 1.5–2.5 × 1.3–1.5 mm, glabrous; glands 2–3, elliptic, 0.6–0.8 × 0.8–1.2 mm; horns absent. |
campanulate, 1.1–1.8 × 1.5–2.5 mm, glabrous; glands 4, yellowish, sessile or short stipitate, circular to oblong, 0.5–0.7 × 0.5–0.8 mm; appendages white to yellowish, ovate to oblong, occasionally rudimentary, (0–)0.8–1.2 × 0.8–1.5 mm, distal margin entire. |
Staminate flowers | 15–40. |
15–25. |
Pistillate flowers | ovary glabrous; styles 1.5–2 mm, 2-fid. |
ovary glabrous; styles 0.7–1 mm, 2-fid 1/2 length. |
Capsules | globose, 3–4.5 × 3–4.5 mm, slightly 3-lobed; cocci rounded, verrucose-tuberculate, glabrous; columella 2.5–3.3 mm. |
ovoid, 4.7–5.5(–6) × 3.6–5.1 mm, glabrous; columella 4.3–5.1 mm. |
Seeds | brown, ovoid, 2.4–2.6 × 1.3–2 mm, smooth, caruncle reniform, 0.2–0.3 × 0.8–0.9 mm. |
grayish white to reddish brown mottled, bottle-shaped, strongly dorsiventrally compressed and weakly 3-angled in cross section, (2.8–)4.1–5.2 × 1.2–2(–3.4) mm, smooth; carunclelike structure linear, 0.4–0.5 × 0.1–0.2 mm. |
Cyathial | arrangement: terminal pleiochasial branches 3–5, each 2–3 times 2–4-branched; pleiochasial bracts ovate, similar in size to distal leaves; dichasial bracts distinct, ovate to suborbiculate, base truncate or rounded, margins entire or finely denticulate, apex obtuse, sometimes mucronulate; axillary cymose branches 0–4. |
|
Cyathia | peduncle 1–5 mm. |
solitary at distal nodes; peduncle 1.4–3.6 mm. |
Euphorbia oblongata |
Euphorbia carunculata |
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Phenology | Flowering and fruiting spring–fall. | Flowering and fruiting summer–fall. |
Habitat | Waste areas, disturbed sites, roadsides, fields, pastures. | Sand dunes. |
Elevation | 30–900 m. (100–3000 ft.) | 400–1300 m. (1300–4300 ft.) |
Distribution |
CA; OR; WA; s Europe [Introduced in North America]
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KS; NM; OK; TX; Mexico (Chihuahua) |
Discussion | Euphorbia oblongata is listed as a noxious weed by the states of California, Oregon, and Washington. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia carunculata has a highly localized and scattered distribution. The species is restricted to sand dunes and known from only a handful of localities throughout its relatively wide range. The seeds are unique in being bottle-shaped and strongly laterally compressed, and unlike other members of sect. Anisophyllum, there is a minute, linear, carunclelike protuberance at the hilum. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 305. | FNA vol. 12, p. 262. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Tithymalus oblongatus | Chamaesyce carunculata |
Name authority | Grisebach: Spic. Fl. Rumel. 1: 136. (1843) | Waterfall: Rhodora 50: 63. (1948) |
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