Euphorbia oblongata |
Euphorbia angusta |
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Balkan spurge, egg-leaf spurge, oblong spurge |
Blackfoot sandmat, narrow-leaf spurge |
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Habit | Herbs, perennial, with woody taproot. | Herbs, perennial, with moderately to strongly thickened rootstock. |
Stems | erect, unbranched or densely branching, 80 cm, often densely villous (especially young stems and pleiochasial branches). |
erect, 12–43 cm, uniformly strigose. |
Leaves | petiole absent; blade oblong to narrowly obovate or lanceolate, 15–70 × 6–25 mm, base rounded or truncate, margins finely serrulate, apex obtuse, mucronulate, surfaces glabrous; venation inconspicuously pinnate, midvein prominent. |
opposite; stipules distinct, linear-subulate or nodiform to papilliform nodiform to papilliform stipules often reddish brown, 0.1–0.7 mm, strigose; petiole 0.3–1.2 mm, strigose; blade: proximal ovate to ovate-elliptic, distal linear to elliptic-linear, 7–41 × 2–5 mm, distal leaf blades more than 6 times as long as wide, base asymmetric, cuneate to rounded, margins entire, often involute on drying, apex acute, surfaces usually short strigose, occasionally glabrous adaxially; venation pinnate, only midvein conspicuous. |
Involucre | cupulate to slightly turbinate, 1.5–2.5 × 1.3–1.5 mm, glabrous; glands 2–3, elliptic, 0.6–0.8 × 0.8–1.2 mm; horns absent. |
turbinate to campanulate-turbinate, 1–1.5 × 1–1.4 mm, strigose; glands 4, green to yellow-green, concave, narrowly oblong, 0.2–0.4 × 0.4–0.7 mm; appendages white, flabellate, 0.5–1.1 × 0.3–0.5 mm, distal margin shallowly and irregularly toothed. |
Staminate flowers | 15–40. |
16–26. |
Pistillate flowers | ovary glabrous; styles 1.5–2 mm, 2-fid. |
ovary strigose; styles 0.3–0.5 mm, 2-fid at apex to almost 1/2 length. |
Capsules | globose, 3–4.5 × 3–4.5 mm, slightly 3-lobed; cocci rounded, verrucose-tuberculate, glabrous; columella 2.5–3.3 mm. |
broadly ovoid, 2–2.6 × 2.5–3 mm, strigose; columella 1.6–2.2 mm. |
Seeds | brown, ovoid, 2.4–2.6 × 1.3–2 mm, smooth, caruncle reniform, 0.2–0.3 × 0.8–0.9 mm. |
white, ovoid, 4-angled in cross section, 1.7–2.2 × 1.1–1.2 mm, transversely low-ridged or wrinkled. |
Cyathial | arrangement: terminal pleiochasial branches 3–5, each 2–3 times 2–4-branched; pleiochasial bracts ovate, similar in size to distal leaves; dichasial bracts distinct, ovate to suborbiculate, base truncate or rounded, margins entire or finely denticulate, apex obtuse, sometimes mucronulate; axillary cymose branches 0–4. |
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Cyathia | peduncle 1–5 mm. |
solitary at distal nodes; peduncle 1.1–2.4 mm. |
2n | = 28. |
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Euphorbia oblongata |
Euphorbia angusta |
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Phenology | Flowering and fruiting spring–fall. | Flowering and fruiting early spring–fall. |
Habitat | Waste areas, disturbed sites, roadsides, fields, pastures. | Rocky limestone soils. |
Elevation | 30–900 m. (100–3000 ft.) | 400–1200 m. (1300–3900 ft.) |
Distribution |
CA; OR; WA; s Europe [Introduced in North America]
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TX; Mexico (Coahuila) |
Discussion | Euphorbia oblongata is listed as a noxious weed by the states of California, Oregon, and Washington. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia angusta, which in the flora area is known from the trans-Pecos region to the Edwards Plateau, is easily recognized by its erect habit, linear leaves, and relatively showy involucral gland appendages with toothed margins. The species is closely related to E. acuta and the Mexican endemic E. johnstonii Mayfield (M. H. Mayfield 1991); it is not only morphologically distinctive but is also the only species in sect. Anisophyllum with C3 photosynthesis (R. F. Sage et al. 2011; T. L. Sage et al. 2011; G. L. Webster 1975). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 305. | FNA vol. 12, p. 260. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Tithymalus oblongatus | Chamaesyce angusta |
Name authority | Grisebach: Spic. Fl. Rumel. 1: 136. (1843) | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 189. (1859) |
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