Euphorbia nutans |
Euphorbia peplus |
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eyebane, eyebane sandmat, eyebane spurge, nodding or upright spotted spurge, nodding spurge |
petty spurge |
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Habit | Herbs, annual, with taproot. | Herbs, annual, with taproot. |
Stems | usually ascending, occasionally erect, often arched at tips, 20–80 cm, sparsely to moderately pilose to villous or with short, incurved hairs, hairs often concentrated at nodes and distally, occasionally in 2 bands along opposite sides of stem. |
erect, unbranched or branched, 5–30 cm, glabrous. |
Leaves | opposite; stipules usually distinct, sometimes connate basally on one side of stem, small scales, irregularly toothed, fringed, or divided, 1–1.5 mm, sparsely to moderately villous distally; petiole 0.3–1.6 mm, moderately pilose to villous; blade oblong to oblong-lanceolate, 8–40 × 3–12 mm, base asymmetric, one side usually angled or rounded, other side ± truncate to cordate-auriculate, margins serrulate, apex angled with blunt tip, abaxial surface pale green or faintly to strongly reddish tinged, adaxial surface usually reddish-mottled or with conspicuous reddish spot, both surfaces usually sparsely to moderately pilose, especially toward base, sometimes glabrous; 3–5-veined from base, pinnate distally, veins faint. |
petiole 1–10 mm; blade obovate, oblong, or suborbiculate, 5–25 × 4–15 mm, base attenuate or cuneate, margins entire, apex obtuse, surfaces glabrous; venation pinnate, midvein prominent. |
Involucre | narrowly obconic, 0.5–1 × 0.3–0.7 mm, glabrous; glands 4, usually green, sometimes reddish purple, oblong to nearly circular, 0.2–0.4 × 0.3–0.5 mm; appendages white or pinkish, ovate to broadly elliptic, 0.2–1 × 0.2–1.5 mm, distal margin entire. |
cupulate to slightly turbinate, 0.6–1.1 × 0.7–0.9 mm, glabrous; glands 4, elliptic, 0.2–0.3 × 0.5–0.6 mm; horns slightly convergent to divergent, 0.4–0.6 mm. |
Staminate flowers | 5–28. |
10–15. |
Pistillate flowers | ovary glabrous; styles 0.6–2.5 mm, 2-fid 1/2 length. |
ovary glabrous; styles 0.1–0.2 mm, 2-fid. |
Capsules | ovoid, 1.6–2.3 × 1.5–2.4 mm, glabrous; columella 1.4–1.6 mm. |
subglobose, 1.3–2 × 1.5–2.2 mm, slightly 3-lobed; cocci rounded, with 2 low longitudinal wings, smooth, glabrous; columella 1.1–1.3 mm. |
Seeds | dark brown, sometimes with thin, white coating (often more persistent along angles than faces), elliptic-ovoid to ovoid, rounded-angular in cross section, 1–1.6 × 0.5–0.8 mm, surface finely and irregularly wrinkled, sometimes faintly so, or with indistinct, shallow, rounded cross ridges. |
whitish or grayish, subovoid, 1–1.6 × 0.6–1 mm, abaxial faces regularly large-pitted (appearing almost alveolate), adaxial faces longitudinally sulcate; caruncle deciduous, conic, 0.2–0.5 × 0.2–0.7 mm. |
Cyathia | solitary at distal nodes or in small, cymose clusters at branch tips; peduncle 0.5–2.5 mm. |
peduncle 0.4–1.1 mm. |
Cyathial | arrangement: terminal pleiochasial branches 3–5, usually 2–8 times 2-branched, 1st branching level rarely 3-furcate; pleiochasial bracts similar in shape to and slightly larger than distal leaves; dichasial bracts distinct, ovate to obovate, base obtuse, margins entire, apex obtuse, usually mucronate; axillary cymose branches 0–3. |
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2n | = 12, 14, 22. |
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Euphorbia nutans |
Euphorbia peplus |
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Phenology | Flowering and fruiting spring–early fall. | Flowering and fruiting year-round. |
Habitat | Stream banks, pond edges, disturbed portions of upland prairies, mesic to dry upland forest openings, pastures, fallow fields, railroads, roadsides, gardens, disturbed areas. | Edges of gardens, weedy flower beds, roadsides, waste places, open ground near forests. |
Elevation | 0–1600 m. (0–5200 ft.) | 0–1500 m. (0–4900 ft.) |
Distribution |
AL; AR; CA; CT; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; MI; MN; MO; MS; NC; ND; NE; NH; NJ; NM; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV; WY; ON; QC; Mexico; Central America; South America; West Indies [Introduced in Eurasia]
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AL; AZ; CA; CO; CT; DE; IA; ID; IL; IN; KY; LA; MA; MD; ME; MI; MT; NC; ND; NH; NJ; NM; NV; NY; OH; OR; PA; RI; TN; TX; UT; VA; VT; WA; WI; WV; AB; BC; MB; NB; NF; NS; ON; PE; QC; SK; Europe; w Asia [Introduced in North America; introduced also in Mexico, West Indies, Bermuda, Central America, South America, Africa, Atlantic Islands, Indian Ocean Islands, Pacific Islands, Australia]
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Discussion | Euphorbia nutans is probably native to at least central and eastern North America, but given its strongly weedy tendencies, it is difficult to know where it may be adventive in parts of the flora area. It is certainly introduced where it occurs in the Old World and probably in South America as well. D. G. Burch (1966) discussed the problems of assigning names to the four main entities in this nomenclatural complex (E. hypericifolia, E. hyssopifolia, E. lasiocarpa, and E. nutans) and determined that the oldest valid name for the relatively robust, temperate North American plants with ascending stems is E. nutans. See the treatment of 56. E. maculata for a discussion of the misapplication of that name to E. nutans. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 12, p. 278. | FNA vol. 12, p. 307. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > subg. Esula |
Sibling taxa | ||
Synonyms | Chamaesyce lansingii, C. nutans, E. lansingii, E. preslii | Esula peplus, Galarhoeus peplus, Tithymalus peplus |
Name authority | Lagasca: Gen. Sp. Pl., 17. (1816) | Linnaeus: Sp. Pl. 1: 456. (1753) |
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