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eyebane, eyebane sandmat, eyebane spurge, nodding or upright spotted spurge, nodding spurge

petty spurge

Habit Herbs, annual, with taproot. Herbs, annual, with taproot.
Stems

usually ascending, occasionally erect, often arched at tips, 20–80 cm, sparsely to moderately pilose to villous or with short, incurved hairs, hairs often concentrated at nodes and distally, occasionally in 2 bands along opposite sides of stem.

erect, unbranched or branched, 5–30 cm, glabrous.

Leaves

opposite;

stipules usually distinct, sometimes connate basally on one side of stem, small scales, irregularly toothed, fringed, or divided, 1–1.5 mm, sparsely to moderately villous distally;

petiole 0.3–1.6 mm, moderately pilose to villous;

blade oblong to oblong-lanceolate, 8–40 × 3–12 mm, base asymmetric, one side usually angled or rounded, other side ± truncate to cordate-auriculate, margins serrulate, apex angled with blunt tip, abaxial surface pale green or faintly to strongly reddish tinged, adaxial surface usually reddish-mottled or with conspicuous reddish spot, both surfaces usually sparsely to moderately pilose, especially toward base, sometimes glabrous; 3–5-veined from base, pinnate distally, veins faint.

petiole 1–10 mm;

blade obovate, oblong, or suborbiculate, 5–25 × 4–15 mm, base attenuate or cuneate, margins entire, apex obtuse, surfaces glabrous;

venation pinnate, midvein prominent.

Involucre

narrowly obconic, 0.5–1 × 0.3–0.7 mm, glabrous;

glands 4, usually green, sometimes reddish purple, oblong to nearly circular, 0.2–0.4 × 0.3–0.5 mm;

appendages white or pinkish, ovate to broadly elliptic, 0.2–1 × 0.2–1.5 mm, distal margin entire.

cupulate to slightly turbinate, 0.6–1.1 × 0.7–0.9 mm, glabrous;

glands 4, elliptic, 0.2–0.3 × 0.5–0.6 mm;

horns slightly convergent to divergent, 0.4–0.6 mm.

Staminate flowers

5–28.

10–15.

Pistillate flowers

ovary glabrous;

styles 0.6–2.5 mm, 2-fid 1/2 length.

ovary glabrous;

styles 0.1–0.2 mm, 2-fid.

Capsules

ovoid, 1.6–2.3 × 1.5–2.4 mm, glabrous;

columella 1.4–1.6 mm.

subglobose, 1.3–2 × 1.5–2.2 mm, slightly 3-lobed;

cocci rounded, with 2 low longitudinal wings, smooth, glabrous;

columella 1.1–1.3 mm.

Seeds

dark brown, sometimes with thin, white coating (often more persistent along angles than faces), elliptic-ovoid to ovoid, rounded-angular in cross section, 1–1.6 × 0.5–0.8 mm, surface finely and irregularly wrinkled, sometimes faintly so, or with indistinct, shallow, rounded cross ridges.

whitish or grayish, subovoid, 1–1.6 × 0.6–1 mm, abaxial faces regularly large-pitted (appearing almost alveolate), adaxial faces longitudinally sulcate;

caruncle deciduous, conic, 0.2–0.5 × 0.2–0.7 mm.

Cyathia

solitary at distal nodes or in small, cymose clusters at branch tips;

peduncle 0.5–2.5 mm.

peduncle 0.4–1.1 mm.

Cyathial

arrangement: terminal pleiochasial branches 3–5, usually 2–8 times 2-branched, 1st branching level rarely 3-furcate;

pleiochasial bracts similar in shape to and slightly larger than distal leaves;

dichasial bracts distinct, ovate to obovate, base obtuse, margins entire, apex obtuse, usually mucronate;

axillary cymose branches 0–3.

2n

= 12, 14, 22.

Euphorbia nutans

Euphorbia peplus

Phenology Flowering and fruiting spring–early fall. Flowering and fruiting year-round.
Habitat Stream banks, pond edges, disturbed portions of upland prairies, mesic to dry upland forest openings, pastures, fallow fields, railroads, roadsides, gardens, disturbed areas. Edges of gardens, weedy flower beds, roadsides, waste places, open ground near forests.
Elevation 0–1600 m. (0–5200 ft.) 0–1500 m. (0–4900 ft.)
Distribution
from FNA
AL; AR; CA; CT; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; MI; MN; MO; MS; NC; ND; NE; NH; NJ; NM; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV; WY; ON; QC; Mexico; Central America; South America; West Indies [Introduced in Eurasia]
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AZ; CA; CO; CT; DE; IA; ID; IL; IN; KY; LA; MA; MD; ME; MI; MT; NC; ND; NH; NJ; NM; NV; NY; OH; OR; PA; RI; TN; TX; UT; VA; VT; WA; WI; WV; AB; BC; MB; NB; NF; NS; ON; PE; QC; SK; Europe; w Asia [Introduced in North America; introduced also in Mexico, West Indies, Bermuda, Central America, South America, Africa, Atlantic Islands, Indian Ocean Islands, Pacific Islands, Australia]
[WildflowerSearch map]
[BONAP county map]
Discussion

Euphorbia nutans is probably native to at least central and eastern North America, but given its strongly weedy tendencies, it is difficult to know where it may be adventive in parts of the flora area. It is certainly introduced where it occurs in the Old World and probably in South America as well. D. G. Burch (1966) discussed the problems of assigning names to the four main entities in this nomenclatural complex (E. hypericifolia, E. hyssopifolia, E. lasiocarpa, and E. nutans) and determined that the oldest valid name for the relatively robust, temperate North American plants with ascending stems is E. nutans. See the treatment of 56. E. maculata for a discussion of the misapplication of that name to E. nutans.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 278. FNA vol. 12, p. 307.
Parent taxa Euphorbiaceae > Euphorbia > sect. Anisophyllum Euphorbiaceae > Euphorbia > subg. Esula
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms Chamaesyce lansingii, C. nutans, E. lansingii, E. preslii Esula peplus, Galarhoeus peplus, Tithymalus peplus
Name authority Lagasca: Gen. Sp. Pl., 17. (1816) Linnaeus: Sp. Pl. 1: 456. (1753)
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