Euphorbia nutans |
Euphorbia longicruris |
|
---|---|---|
eyebane, eyebane sandmat, eyebane spurge, nodding or upright spotted spurge, nodding spurge |
wedge-leaf spurge |
|
Habit | Herbs, annual, with taproot. | Herbs, annual, with taproot. |
Stems | usually ascending, occasionally erect, often arched at tips, 20–80 cm, sparsely to moderately pilose to villous or with short, incurved hairs, hairs often concentrated at nodes and distally, occasionally in 2 bands along opposite sides of stem. |
erect, usually unbranched, occasionally branched later in season, 5–25 cm, glabrous. |
Leaves | opposite; stipules usually distinct, sometimes connate basally on one side of stem, small scales, irregularly toothed, fringed, or divided, 1–1.5 mm, sparsely to moderately villous distally; petiole 0.3–1.6 mm, moderately pilose to villous; blade oblong to oblong-lanceolate, 8–40 × 3–12 mm, base asymmetric, one side usually angled or rounded, other side ± truncate to cordate-auriculate, margins serrulate, apex angled with blunt tip, abaxial surface pale green or faintly to strongly reddish tinged, adaxial surface usually reddish-mottled or with conspicuous reddish spot, both surfaces usually sparsely to moderately pilose, especially toward base, sometimes glabrous; 3–5-veined from base, pinnate distally, veins faint. |
petiole 0–0.5 mm; blade cuneate-spatulate to obovate, 5–15 × 2–6 mm, base broadly attenuate, margins entire, apex rounded to obtuse, mucronate, surfaces glabrous; venation pinnate, midvein prominent. |
Involucre | narrowly obconic, 0.5–1 × 0.3–0.7 mm, glabrous; glands 4, usually green, sometimes reddish purple, oblong to nearly circular, 0.2–0.4 × 0.3–0.5 mm; appendages white or pinkish, ovate to broadly elliptic, 0.2–1 × 0.2–1.5 mm, distal margin entire. |
campanulate, 1.5–2 × 1–1.5 mm, glabrous; glands 4, crescent-shaped to elliptic, 0.4–0.8 × 0.8–1.1 mm; horns divergent, 0.5–0.8 mm. |
Staminate flowers | 5–28. |
10–15. |
Pistillate flowers | ovary glabrous; styles 0.6–2.5 mm, 2-fid 1/2 length. |
ovary glabrous; styles 0.5–0.6 mm, 2-fid. |
Capsules | ovoid, 1.6–2.3 × 1.5–2.4 mm, glabrous; columella 1.4–1.6 mm. |
ovoid-globose, 2–2.8 × 2.5–3 mm, 3-lobed; cocci rounded, smooth, glabrous; columella 1.6–2.1 mm. |
Seeds | dark brown, sometimes with thin, white coating (often more persistent along angles than faces), elliptic-ovoid to ovoid, rounded-angular in cross section, 1–1.6 × 0.5–0.8 mm, surface finely and irregularly wrinkled, sometimes faintly so, or with indistinct, shallow, rounded cross ridges. |
gray to purple-gray or sometimes nearly black, oblong, 1.3–1.6 × 0.9–1.2 mm, strongly small-pitted; caruncle umbonate, depressed-conic, 0.5 × 0.7 mm. |
Cyathia | solitary at distal nodes or in small, cymose clusters at branch tips; peduncle 0.5–2.5 mm. |
peduncle 0.3–0.5 mm. |
Cyathial | arrangement: terminal pleiochasial branches 3, each many times 2-branched; pleiochasial bracts obovate, similar in size to distal leaves; dichasial bracts basally subconnate, strongly imbricate and often obscuring internodes, reniform to semiorbiculate, base cordate, margins entire, apex rounded; axillary cymose branches 0–5. |
|
2n | = 12, 14, 22. |
|
Euphorbia nutans |
Euphorbia longicruris |
|
Phenology | Flowering and fruiting spring–early fall. | Flowering and fruiting spring. |
Habitat | Stream banks, pond edges, disturbed portions of upland prairies, mesic to dry upland forest openings, pastures, fallow fields, railroads, roadsides, gardens, disturbed areas. | Grasslands, open prairies, sites with rocky, usually calcareous soils. |
Elevation | 0–1600 m. (0–5200 ft.) | 300–800 m. (1000–2600 ft.) |
Distribution |
AL; AR; CA; CT; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; MI; MN; MO; MS; NC; ND; NE; NH; NJ; NM; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV; WY; ON; QC; Mexico; Central America; South America; West Indies [Introduced in Eurasia]
|
AR; KS; OK; TX |
Discussion | Euphorbia nutans is probably native to at least central and eastern North America, but given its strongly weedy tendencies, it is difficult to know where it may be adventive in parts of the flora area. It is certainly introduced where it occurs in the Old World and probably in South America as well. D. G. Burch (1966) discussed the problems of assigning names to the four main entities in this nomenclatural complex (E. hypericifolia, E. hyssopifolia, E. lasiocarpa, and E. nutans) and determined that the oldest valid name for the relatively robust, temperate North American plants with ascending stems is E. nutans. See the treatment of 56. E. maculata for a discussion of the misapplication of that name to E. nutans. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia longicruris is quite similar to the other small, annual members of subg. Esula in the south-central United States and can best be distinguished from those species by its imbricate dichasial bracts that form little tufts of overlapping leaves at the ends of the pleiochasial branches. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 278. | FNA vol. 12, p. 304. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > subg. Esula |
Sibling taxa | ||
Synonyms | Chamaesyce lansingii, C. nutans, E. lansingii, E. preslii | Tithymalus longicruris |
Name authority | Lagasca: Gen. Sp. Pl., 17. (1816) | Scheele: Linnaea 22: 152. (1849) |
Web links |
|