Euphorbia nutans |
Euphorbia hyssopifolia |
|
---|---|---|
eyebane, eyebane sandmat, eyebane spurge, nodding or upright spotted spurge, nodding spurge |
hyssopleaf sandmat |
|
Habit | Herbs, annual, with taproot. | Herbs, annual, with taproot. |
Stems | usually ascending, occasionally erect, often arched at tips, 20–80 cm, sparsely to moderately pilose to villous or with short, incurved hairs, hairs often concentrated at nodes and distally, occasionally in 2 bands along opposite sides of stem. |
erect to ascending, 80 cm, sparsely to densely pilose or pilose-crinkled proximally, usually glabrous distally. |
Leaves | opposite; stipules usually distinct, sometimes connate basally on one side of stem, small scales, irregularly toothed, fringed, or divided, 1–1.5 mm, sparsely to moderately villous distally; petiole 0.3–1.6 mm, moderately pilose to villous; blade oblong to oblong-lanceolate, 8–40 × 3–12 mm, base asymmetric, one side usually angled or rounded, other side ± truncate to cordate-auriculate, margins serrulate, apex angled with blunt tip, abaxial surface pale green or faintly to strongly reddish tinged, adaxial surface usually reddish-mottled or with conspicuous reddish spot, both surfaces usually sparsely to moderately pilose, especially toward base, sometimes glabrous; 3–5-veined from base, pinnate distally, veins faint. |
opposite; stipules usually connate, irregularly lacerate, 0.5–1 mm, usually glabrous, occasionally with few marginal hairs; petiole 1–2 mm, glabrous; blade lanceolate to oblong or falcate, 8–35 × 7–15 mm, base asymmetric, rounded, margins serrulate, apex broadly acute, abaxial surface glabrous or sparsely pilose toward base, adaxial surface glabrous; palmately veined at base, pinnate distally. |
Involucre | narrowly obconic, 0.5–1 × 0.3–0.7 mm, glabrous; glands 4, usually green, sometimes reddish purple, oblong to nearly circular, 0.2–0.4 × 0.3–0.5 mm; appendages white or pinkish, ovate to broadly elliptic, 0.2–1 × 0.2–1.5 mm, distal margin entire. |
obconic, 0.9–1.1 × 0.7–0.9 mm, glabrous; glands 4(–5) (5th gland without appendage), yellow-green to maroon, elliptic to circular, 0.1–0.2 × 0.1–0.3 mm; appendages spreading, usually white or turning reddish with age, short reniform or semilunate, 0.1–0.3 × 0.2–0.6 mm, distal margin entire or slightly undulate to crenate. |
Staminate flowers | 5–28. |
4–15. |
Pistillate flowers | ovary glabrous; styles 0.6–2.5 mm, 2-fid 1/2 length. |
ovary glabrous; styles 0.5–0.9 mm, 2-fid 1/2 length. |
Capsules | ovoid, 1.6–2.3 × 1.5–2.4 mm, glabrous; columella 1.4–1.6 mm. |
depressed-ovoid, 1.5–1.6 × 1.7–1.8 mm, glabrous; columella 1.5–2 mm. |
Seeds | dark brown, sometimes with thin, white coating (often more persistent along angles than faces), elliptic-ovoid to ovoid, rounded-angular in cross section, 1–1.6 × 0.5–0.8 mm, surface finely and irregularly wrinkled, sometimes faintly so, or with indistinct, shallow, rounded cross ridges. |
brown to grayish white, ovoid, slightly 4-angled in cross section, abaxial faces convex, adaxial faces slightly concave to slightly convex, 1–1.4 × 0.7–1.1 mm, with 2–3 prominent transverse ridges that do not interrupt adaxial keel, or coarsely and inconspicuously pitted-reticulate. |
Cyathia | solitary at distal nodes or in small, cymose clusters at branch tips; peduncle 0.5–2.5 mm. |
solitary or in small, cymose clusters occasionally with bractlike leaves at distal nodes or on congested, axillary branches; peduncle 0.5–2.5 mm. |
2n | = 12, 14, 22. |
= 12, 14. |
Euphorbia nutans |
Euphorbia hyssopifolia |
|
Phenology | Flowering and fruiting spring–early fall. | Flowering and fruiting late spring–early fall. |
Habitat | Stream banks, pond edges, disturbed portions of upland prairies, mesic to dry upland forest openings, pastures, fallow fields, railroads, roadsides, gardens, disturbed areas. | Disturbed areas, ditches, gardens. |
Elevation | 0–1600 m. (0–5200 ft.) | 0–1500 m. (0–4900 ft.) |
Distribution |
AL; AR; CA; CT; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; MI; MN; MO; MS; NC; ND; NE; NH; NJ; NM; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV; WY; ON; QC; Mexico; Central America; South America; West Indies [Introduced in Eurasia]
|
AL; AZ; FL; GA; LA; MS; NM; SC; TX; UT; Mexico; Central America; South America; West Indies [Introduced in tropical Asia, Africa, Australia]
|
Discussion | Euphorbia nutans is probably native to at least central and eastern North America, but given its strongly weedy tendencies, it is difficult to know where it may be adventive in parts of the flora area. It is certainly introduced where it occurs in the Old World and probably in South America as well. D. G. Burch (1966) discussed the problems of assigning names to the four main entities in this nomenclatural complex (E. hypericifolia, E. hyssopifolia, E. lasiocarpa, and E. nutans) and determined that the oldest valid name for the relatively robust, temperate North American plants with ascending stems is E. nutans. See the treatment of 56. E. maculata for a discussion of the misapplication of that name to E. nutans. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia hyssopifolia is native to the New World tropics and is probably also native to parts of the southern United States. However, at least some of the records from the flora area appear to be from adventive plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 278. | FNA vol. 12, p. 272. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Chamaesyce lansingii, C. nutans, E. lansingii, E. preslii | Chamaesyce hyssopifolia, E. jonesii, E. stenomeres |
Name authority | Lagasca: Gen. Sp. Pl., 17. (1816) | Linnaeus: Syst. Nat. ed. 10, 2: 1048. (1759) |
Web links |
|