Euphorbia nutans |
Euphorbia curtisii |
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eyebane, eyebane sandmat, eyebane spurge, nodding or upright spotted spurge, nodding spurge |
Curtis' spurge, Sandhills spurge |
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Habit | Herbs, annual, with taproot. | Herbs, perennial, with spreading rootstock. |
Stems | usually ascending, occasionally erect, often arched at tips, 20–80 cm, sparsely to moderately pilose to villous or with short, incurved hairs, hairs often concentrated at nodes and distally, occasionally in 2 bands along opposite sides of stem. |
erect or ascending, branched, solitary or few, previous year's dead stems not persistent, 20–40 cm, usually glabrous, rarely strigose to sericeous at nodes. |
Leaves | opposite; stipules usually distinct, sometimes connate basally on one side of stem, small scales, irregularly toothed, fringed, or divided, 1–1.5 mm, sparsely to moderately villous distally; petiole 0.3–1.6 mm, moderately pilose to villous; blade oblong to oblong-lanceolate, 8–40 × 3–12 mm, base asymmetric, one side usually angled or rounded, other side ± truncate to cordate-auriculate, margins serrulate, apex angled with blunt tip, abaxial surface pale green or faintly to strongly reddish tinged, adaxial surface usually reddish-mottled or with conspicuous reddish spot, both surfaces usually sparsely to moderately pilose, especially toward base, sometimes glabrous; 3–5-veined from base, pinnate distally, veins faint. |
alternate; stipules to 0.1 mm; petiole to (0–)1–2 mm, glabrous or strigose to sericeous; blade usually linear, occasionally elliptic, rarely ovate, proximal often greatly reduced and often scalelike, 10–30 × 1.5–6 mm, base cuneate, margins entire, occasionally sparsely ciliate, apex rounded or broadly acute, abaxial surface glabrous or sparsely strigose to sericeous, adaxial surface glabrous; venation obscure, only midvein conspicuous. |
Involucre | narrowly obconic, 0.5–1 × 0.3–0.7 mm, glabrous; glands 4, usually green, sometimes reddish purple, oblong to nearly circular, 0.2–0.4 × 0.3–0.5 mm; appendages white or pinkish, ovate to broadly elliptic, 0.2–1 × 0.2–1.5 mm, distal margin entire. |
campanulate, 1–1.2 × 1.3–1.5(–1.7) mm, glabrous or strigose to sericeous on distal 1/2; glands 5, green, reniform, 0.3 × 0.6 mm; appendages white, semicircular, 0.3–0.4 × 0.6–0.8 mm, entire. |
Staminate flowers | 5–28. |
20–25. |
Pistillate flowers | ovary glabrous; styles 0.6–2.5 mm, 2-fid 1/2 length. |
ovary glabrous or sparsely strigose to sericeous; styles 0.6–1.1 mm, 2-fid at apex to 1/2 length. |
Capsules | ovoid, 1.6–2.3 × 1.5–2.4 mm, glabrous; columella 1.4–1.6 mm. |
globose, 2.5–3.2 × 4.3–5.1 mm, glabrous or sparsely strigose to sericeous; columella 2.4–3.1 mm. |
Seeds | dark brown, sometimes with thin, white coating (often more persistent along angles than faces), elliptic-ovoid to ovoid, rounded-angular in cross section, 1–1.6 × 0.5–0.8 mm, surface finely and irregularly wrinkled, sometimes faintly so, or with indistinct, shallow, rounded cross ridges. |
usually gray to black, occasionally brown, ovoid-globose, 2.2 × 1.8 mm, smooth; caruncle absent. |
Cyathia | solitary at distal nodes or in small, cymose clusters at branch tips; peduncle 0.5–2.5 mm. |
in terminal pleiochasia (fertile axillary branches occasionally present); peduncle 6.5–17 mm, filiform, glabrous. |
2n | = 12, 14, 22. |
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Euphorbia nutans |
Euphorbia curtisii |
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Phenology | Flowering and fruiting spring–early fall. | Flowering and fruiting early spring–summer. |
Habitat | Stream banks, pond edges, disturbed portions of upland prairies, mesic to dry upland forest openings, pastures, fallow fields, railroads, roadsides, gardens, disturbed areas. | Xeric to dry oak or oak-pine scrub of sand hills, pine-oak woodlands, pine-oak savannas. |
Elevation | 0–1600 m. (0–5200 ft.) | 0–200 m. (0–700 ft.) |
Distribution |
AL; AR; CA; CT; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; MI; MN; MO; MS; NC; ND; NE; NH; NJ; NM; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV; WY; ON; QC; Mexico; Central America; South America; West Indies [Introduced in Eurasia]
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FL; GA; NC; SC
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Discussion | Euphorbia nutans is probably native to at least central and eastern North America, but given its strongly weedy tendencies, it is difficult to know where it may be adventive in parts of the flora area. It is certainly introduced where it occurs in the Old World and probably in South America as well. D. G. Burch (1966) discussed the problems of assigning names to the four main entities in this nomenclatural complex (E. hypericifolia, E. hyssopifolia, E. lasiocarpa, and E. nutans) and determined that the oldest valid name for the relatively robust, temperate North American plants with ascending stems is E. nutans. See the treatment of 56. E. maculata for a discussion of the misapplication of that name to E. nutans. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia curtisii is found in the Gulf and Atlantic coastal plains. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 278. | FNA vol. 12, p. 245. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Alectoroctonum |
Sibling taxa | ||
Synonyms | Chamaesyce lansingii, C. nutans, E. lansingii, E. preslii | E. eriogonoides, Tithymalopsis curtisii, T. eriogonoides |
Name authority | Lagasca: Gen. Sp. Pl., 17. (1816) | Engelmann ex Chapman: Fl. South. U.S., 401. (1860) |
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