Euphorbia nutans |
Euphorbia antisyphilitica |
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eyebane, eyebane sandmat, eyebane spurge, nodding or upright spotted spurge, nodding spurge |
candelilla, wax plant |
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Habit | Herbs, annual, with taproot. | Shrubs, with much-branched, fleshy rootstock. |
Stems | usually ascending, occasionally erect, often arched at tips, 20–80 cm, sparsely to moderately pilose to villous or with short, incurved hairs, hairs often concentrated at nodes and distally, occasionally in 2 bands along opposite sides of stem. |
erect, few branched, 25–50(–100) cm, glabrous or puberulent, pencil-like, in age covered with flaky, exfoliating layer of wax. |
Leaves | opposite; stipules usually distinct, sometimes connate basally on one side of stem, small scales, irregularly toothed, fringed, or divided, 1–1.5 mm, sparsely to moderately villous distally; petiole 0.3–1.6 mm, moderately pilose to villous; blade oblong to oblong-lanceolate, 8–40 × 3–12 mm, base asymmetric, one side usually angled or rounded, other side ± truncate to cordate-auriculate, margins serrulate, apex angled with blunt tip, abaxial surface pale green or faintly to strongly reddish tinged, adaxial surface usually reddish-mottled or with conspicuous reddish spot, both surfaces usually sparsely to moderately pilose, especially toward base, sometimes glabrous; 3–5-veined from base, pinnate distally, veins faint. |
alternate usually caducous, sometimes persisting; stipules 0.4–0.5 mm; petiole absent; blade ovate to deltate-subulate, 2.5–4 × 1 mm, thick, fleshy, base usually rounded and swollen, rarely cuneate, margins entire, apex acute, surfaces puberulent, adaxial sometimes canescent; venation inconspicuous. |
Involucre | narrowly obconic, 0.5–1 × 0.3–0.7 mm, glabrous; glands 4, usually green, sometimes reddish purple, oblong to nearly circular, 0.2–0.4 × 0.3–0.5 mm; appendages white or pinkish, ovate to broadly elliptic, 0.2–1 × 0.2–1.5 mm, distal margin entire. |
campanulate, 1.6–2.2 × 1.6–1.9 mm, puberulent to canescent; glands 5, pinkish, narrowly oblong to reniform, 0.3–0.4 × 0.8–1 mm; appendages white to pink, ovate, oblong, or transversely oblong, 1.3–2.5 × 1.4–2.5 mm, usually erose, rarely entire. |
Staminate flowers | 5–28. |
50–70. |
Pistillate flowers | ovary glabrous; styles 0.6–2.5 mm, 2-fid 1/2 length. |
ovary glabrous; styles 0.9–1.1 mm, 2-fid nearly entire length. |
Capsules | ovoid, 1.6–2.3 × 1.5–2.4 mm, glabrous; columella 1.4–1.6 mm. |
oblong to ovoid, 3.9–4.2 × 3.6–3.9 mm, glabrous; columella 3.1–3.3 mm. |
Seeds | dark brown, sometimes with thin, white coating (often more persistent along angles than faces), elliptic-ovoid to ovoid, rounded-angular in cross section, 1–1.6 × 0.5–0.8 mm, surface finely and irregularly wrinkled, sometimes faintly so, or with indistinct, shallow, rounded cross ridges. |
whitish gray, narrowly ovoid, 2.4–3.1 × 1.4–1.6 mm, irregularly rugose-tuberculate; caruncle crescent-shaped, 0.3–0.6 × 0.6–0.8 mm. |
Cyathia | solitary at distal nodes or in small, cymose clusters at branch tips; peduncle 0.5–2.5 mm. |
in axillary congested cymes, near branch tips or solitary at distal nodes; peduncle 0–1 mm, lanulose. |
2n | = 12, 14, 22. |
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Euphorbia nutans |
Euphorbia antisyphilitica |
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Phenology | Flowering and fruiting spring–early fall. | Flowering and fruiting year-round in response to sufficient rainfall. |
Habitat | Stream banks, pond edges, disturbed portions of upland prairies, mesic to dry upland forest openings, pastures, fallow fields, railroads, roadsides, gardens, disturbed areas. | Desert scrub, frequently on limestone substrates. |
Elevation | 0–1600 m. (0–5200 ft.) | 100–1200 m. (300–3900 ft.) |
Distribution |
AL; AR; CA; CT; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; MI; MN; MO; MS; NC; ND; NE; NH; NJ; NM; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV; WY; ON; QC; Mexico; Central America; South America; West Indies [Introduced in Eurasia]
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NM; TX; Mexico
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Discussion | Euphorbia nutans is probably native to at least central and eastern North America, but given its strongly weedy tendencies, it is difficult to know where it may be adventive in parts of the flora area. It is certainly introduced where it occurs in the Old World and probably in South America as well. D. G. Burch (1966) discussed the problems of assigning names to the four main entities in this nomenclatural complex (E. hypericifolia, E. hyssopifolia, E. lasiocarpa, and E. nutans) and determined that the oldest valid name for the relatively robust, temperate North American plants with ascending stems is E. nutans. See the treatment of 56. E. maculata for a discussion of the misapplication of that name to E. nutans. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia antisyphilitica is the only pencil-stemmed species of Euphorbia occurring in the flora area. The species is characteristic of the Chihuahuan Desert scrub of Mexico from Chihuahua and Coahuila south to Hidalgo and Querétaro, and barely enters into the United States in southern New Mexico (Doña Ana and Lincoln counties) and southwest (Brewster, Hudspeth, Presidio, and Terrell counties) and south (Starr and Webb counties) Texas. The stems are covered in a conspicuous coat of exfoliating wax, and the plants historically have been harvested for this product, although the practice is much less prevalent now. The specific epithet refers to its traditional medicinal use in treating sexually transmitted infections. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 278. | FNA vol. 12, p. 243. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Alectoroctonum |
Sibling taxa | ||
Synonyms | Chamaesyce lansingii, C. nutans, E. lansingii, E. preslii | |
Name authority | Lagasca: Gen. Sp. Pl., 17. (1816) | Zuccarini: Flora 15(2, Beibl.): 58. (1832) |
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