Euphorbia myrsinites |
Euphorbia exserta |
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broad leaf glaucous spurge, donkey tail, myrtle or creeping or blue spurge, myrtle spurge |
coastal sand spurge, maroon or purple sand spurge |
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Habit | Herbs, usually perennial, occasionally biennial, with taproot. | Herbs, perennial, with spreading rootstock. |
Stems | erect or semiprostrate, unbranched or branched, 15–40 cm, succulent, glabrous. |
erect or ascending, unbranched or branched, solitary, few, or occasionally densely clumped, previous year's dead stems not persistent, 20–33 cm, usually glabrous, rarely sparsely villous, glaucescent. |
Leaves | petiole 0–2 mm; blade obovate, obovate-oblong, lanceolate, orbiculate, or suborbiculate, 2–30 × 3–17 mm, fleshy, base truncate or attenuate, margins entire or finely denticulate, apex acute to obtuse, cuspidate or strongly mucronate, surfaces glabrous; venation and midvein inconspicuous. |
alternate; stipules less than 0.1 mm; petiole (0–)1–3 mm, glabrous; blade linear or linear-elliptic to obovate or orbiculate, proximal greatly reduced, scalelike, 15–30 × 1–20 mm, base cuneate to rounded, margins entire, apex rounded, broadly acute, or emarginate, surfaces glabrous, glaucescent; venation often obscure on narrow leaves, midvein conspicuous. |
Involucre | campanulate, 2.4–2.6 × 2.3–2.5 mm, glabrous; glands 4, trapezoidal, 1–1.5 × 1.5–2.5 mm; horns divergent, thick, tips rounded, dilated, 0.5–0.9 mm. |
usually dark red, campanulate, 1.3–1.6 × 1.4–2.1 mm, glabrous; glands 5, usually dark red, rarely greenish red, elliptic reniform, thickened, 0.3–0.5 × 0.8 mm; appendages white or green, often forming narrow rim around distal margin of gland, 0–0.2 mm, entire. |
Staminate flowers | 6–12. |
20–25. |
Pistillate flowers | ovary glabrous; styles 2.5–2.8 mm, usually unbranched. |
ovary glabrous; styles 0.4–0.8 mm, 2-fid at apex. |
Capsules | subglobose, 5–7 × 5–6 mm, unlobed; cocci rounded to subangular, smooth, glabrous; columella 4.5–5 mm. |
depressed-globose, 1.8–2.5 × 3.6–4.4 mm, glabrous; columella 1.9–2.4 mm. |
Seeds | brownish to grayish, oblong, 2.8–4.5 × 2–3.2 mm, vermiculate-rugose; caruncle substipitate, trapezoidal or mushroom-shaped, 1.3–1.5 × 0.6–0.8 mm. |
ashy white, ovoid, 2.1 × 1.3 mm, angled with 5 blunt longitudinal ridges, with shallow and irregular pits; caruncle absent. |
Cyathial | arrangement: terminal pleiochasial branches 2–12, each 1–2 times 2-branched; pleiochasial bracts similar in shape and size to distal leaves; dichasial bracts distinct, suborbiculate or reniform, base truncate, margins entire or minutely denticulate, apex obtuse, mucronulate; axillary cymose branches 0–4. |
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Cyathia | peduncle 0.5–1 mm. |
usually in terminal dichasia, sometimes pleiochasia; peduncle 6–33 mm, filiform, glabrous. |
Euphorbia myrsinites |
Euphorbia exserta |
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Phenology | Flowering and fruiting spring–summer. | Flowering and fruiting spring–summer. |
Habitat | Scrub oak communities, open ground near forests, shrub-steppes. | Xeric to dry pine-oak scrub of sand hills, pine-oak woodlands, pine-oak savannas. |
Elevation | 0–2400 m. (0–7900 ft.) | 0–150 m. (0–500 ft.) |
Distribution |
CA; CO; ID; MT; NM; OR; UT; WA; WY; BC; s Europe; w Asia [Introduced in North America]
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FL; GA; NC; SC; VA
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Discussion | Euphorbia myrsinites is cultivated in much of the flora area, where it can tolerate cold winters. In some areas, it can locally escape from cultivation. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
K. R. Park (1998) recognized both Euphorbia exserta and E. gracilior as distinct species, with the former known only from the holotype. However, this treatment follows M. J. Huft (1997) and treats E. gracilior as a synonym of E. exserta. Although the type of E. exserta is unusual in having greenish red (versus dark red) cyathia and glands, as well as small gland appendages, it is otherwise typical of the species as a whole, including features such as the upright habit, reddish coloration, scalelike proximal leaves, and filiform peduncles to 30 mm that are also common to plants formerly treated as E. gracilior. Euphorbia gracilis Elliott, which has sometimes been applied to E. exserta, is an illegitimate name (a later homonym of E. gracilis Loiseleur-Deslongchamps) and pertains here. The upright habit and usually dark red cyathia and glands distinguish Euphorbia exserta from the otherwise similar E. ipecacuanhae, while the glaucescent vegetative parts and smaller gland appendages readily separate E. exserta from the similar E. curtisii. Euphorbia exserta is found on the Gulf and Atlantic coastal plains. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 305. | FNA vol. 12, p. 246. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Alectoroctonum |
Sibling taxa | ||
Synonyms | Tithymalus myrsinites | Tithymalopsis exserta, E. gracilior, T. gracilis |
Name authority | Linnaeus: Sp. Pl. 1: 461. (1753) | (Small) Coker: Pl. Life Hartsville, 88. (1912) |
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