Euphorbia myrsinites |
Euphorbia capitellata |
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broad leaf glaucous spurge, donkey tail, myrtle or creeping or blue spurge, myrtle spurge |
capitate sandmat, head sandmat, head spurge |
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Habit | Herbs, usually perennial, occasionally biennial, with taproot. | Herbs, annual or perennial, with slender to thick and woody rootstock. |
Stems | erect or semiprostrate, unbranched or branched, 15–40 cm, succulent, glabrous. |
usually ascending (but ranging from decumbent to erect), 15–50 cm, glabrous, strigillose, or pilose. |
Leaves | petiole 0–2 mm; blade obovate, obovate-oblong, lanceolate, orbiculate, or suborbiculate, 2–30 × 3–17 mm, fleshy, base truncate or attenuate, margins entire or finely denticulate, apex acute to obtuse, cuspidate or strongly mucronate, surfaces glabrous; venation and midvein inconspicuous. |
opposite; stipules distinct, filiform or divided into 2–3 subulate-filiform segments, without dark, circular glands at base, 0.6–1.5 mm, pilose; petioles 0.6–1.3 mm, glabrous, pilose, or strigillose; blade ovate to narrowly ovate, 4–19 × 2–8 mm, base asymmetric, one side strongly cordate, other side rounded to slightly cordate, margins entire or serrulate (commonly nearly entire with few scattered teeth, often slightly thickened), apex acute, surfaces often with red spot in center, glabrous, pilose, or strigillose; weakly 3-veined from base, usually only midvein conspicuous. |
Involucre | campanulate, 2.4–2.6 × 2.3–2.5 mm, glabrous; glands 4, trapezoidal, 1–1.5 × 1.5–2.5 mm; horns divergent, thick, tips rounded, dilated, 0.5–0.9 mm. |
narrowly obconic to narrowly campanulate, 0.8–1.6 × 0.7–1.3 mm, glabrous or pilose; glands 4, yellow-green to pink or maroon, circular to oblong, 0.2–0.4 × 0.2–0.5 mm; appendages white to light pink, oblong to reniform or flabellate, 0.2–1.1 × 0.5–1.7 mm, surfaces glabrous, distal margin entire. |
Staminate flowers | 6–12. |
25–40. |
Pistillate flowers | ovary glabrous; styles 2.5–2.8 mm, usually unbranched. |
ovary glabrous or pilose; styles 0.4–0.6 mm, 2-fid entire length. |
Capsules | subglobose, 5–7 × 5–6 mm, unlobed; cocci rounded to subangular, smooth, glabrous; columella 4.5–5 mm. |
ovoid to oblate, 1.3–1.9 × 1.4–2.1 mm, glabrous or pilose; columella 1.1–1.7 mm. |
Seeds | brownish to grayish, oblong, 2.8–4.5 × 2–3.2 mm, vermiculate-rugose; caruncle substipitate, trapezoidal or mushroom-shaped, 1.3–1.5 × 0.6–0.8 mm. |
pink to pinkish gray, narrowly ovoid to narrowly ovoid-oblong, 4-angled or weakly 3-angled in cross section, 0.9–1.5 × 0.5–0.7 mm, irregularly dimpled, sometimes also with faint transverse ridges that do not pass through abaxial keel. |
Cyathial | arrangement: terminal pleiochasial branches 2–12, each 1–2 times 2-branched; pleiochasial bracts similar in shape and size to distal leaves; dichasial bracts distinct, suborbiculate or reniform, base truncate, margins entire or minutely denticulate, apex obtuse, mucronulate; axillary cymose branches 0–4. |
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Cyathia | peduncle 0.5–1 mm. |
in dense, terminal capitate glomerules, with reduced, bractlike leaves subtending cyathia, at tips of main stems and short, leafy, axillary branches; peduncle 0.1–1.2 mm. |
2n | = 14. |
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Euphorbia myrsinites |
Euphorbia capitellata |
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Phenology | Flowering and fruiting spring–summer. | Flowering and fruiting year-round in response to sufficient moisture. |
Habitat | Scrub oak communities, open ground near forests, shrub-steppes. | Gravelly washes, rocky slopes, basaltic talus, disturbed roadsides, primarily desert scrub, desert grasslands, riparian forests, rarely oak-juniper woodlands. |
Elevation | 0–2400 m. (0–7900 ft.) | 600–1600 m. (2000–5200 ft.) |
Distribution |
CA; CO; ID; MT; NM; OR; UT; WA; WY; BC; s Europe; w Asia [Introduced in North America]
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AZ; NM; TX; Mexico (Baja California, Baja California Sur, Chihuahua, Coahuila, Sinaloa, Sonora)
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Discussion | Euphorbia myrsinites is cultivated in much of the flora area, where it can tolerate cold winters. In some areas, it can locally escape from cultivation. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia capitellata is a characteristic herb in the Sonoran Desert of southern Arizona, ranging east to extreme southwestern Texas. During peak flowering, plants are attractive due to the dense clusters of cyathia with well-developed involucral gland appendages. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 305. | FNA vol. 12, p. 262. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Tithymalus myrsinites | Chamaesyce capitellata, C. pycnanthema, E. pycnanthema |
Name authority | Linnaeus: Sp. Pl. 1: 461. (1753) | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 188. (1859) |
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