Euphorbia myrsinites |
Euphorbia astyla |
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broad leaf glaucous spurge, donkey tail, myrtle or creeping or blue spurge, myrtle spurge |
alkali spurge, Pecos spurge |
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Habit | Herbs, usually perennial, occasionally biennial, with taproot. | Herbs, perennial, with woody or fibrous-fleshy taproot, 5–12 mm thick. |
Stems | erect or semiprostrate, unbranched or branched, 15–40 cm, succulent, glabrous. |
decumbent, ascending, or erect, few to many emerging from woody crown, 5–25(–50) cm, glabrous. |
Leaves | petiole 0–2 mm; blade obovate, obovate-oblong, lanceolate, orbiculate, or suborbiculate, 2–30 × 3–17 mm, fleshy, base truncate or attenuate, margins entire or finely denticulate, apex acute to obtuse, cuspidate or strongly mucronate, surfaces glabrous; venation and midvein inconspicuous. |
opposite; stipules connate into deltate scale, 0.2–0.5 mm, minutely lacerate at apex, glabrous; petiole 0–0.2(–0.3) mm, glabrous; blade orbiculate-reniform to acute-cordate, 2–5(–8) × 2–5(–6) mm, base ± asymmetric, cordate to auriculate, sometimes clasping stem, margins entire, apex narrowly acute, surfaces glabrous; 2- or 3-veined from base, but usually only midvein conspicuous. |
Involucre | campanulate, 2.4–2.6 × 2.3–2.5 mm, glabrous; glands 4, trapezoidal, 1–1.5 × 1.5–2.5 mm; horns divergent, thick, tips rounded, dilated, 0.5–0.9 mm. |
broadly campanulate, 0.8–1.4 × 0.9–1.4 mm, glabrous; glands 4, yellow-green to brownish, oblong, 0.2–0.3 × 0.5–0.7 mm; appendages white, flabellate to oblong, 0.1–0.2(–0.5) × 0.4–0.8 mm, distal margin entire or dentate-crenate. |
Staminate flowers | 6–12. |
22–26. |
Pistillate flowers | ovary glabrous; styles 2.5–2.8 mm, usually unbranched. |
ovary glabrous; styles 0.3–0.4 mm, unbranched, thickened-clavate. |
Capsules | subglobose, 5–7 × 5–6 mm, unlobed; cocci rounded to subangular, smooth, glabrous; columella 4.5–5 mm. |
ovoid and broadly triangular, 1.5–1.9(–2.5) × 1.4–1.6(–2.2) mm, glabrous; columella 1.2–1.8 mm. |
Seeds | brownish to grayish, oblong, 2.8–4.5 × 2–3.2 mm, vermiculate-rugose; caruncle substipitate, trapezoidal or mushroom-shaped, 1.3–1.5 × 0.6–0.8 mm. |
white, oblong, 4-angled in cross section, adaxial faces slightly concave, with long raphe between, 1.5–1.8 × 0.7–1 mm, markedly foveolate, with irregular to ± parallel or anastomosing ridges. |
Cyathial | arrangement: terminal pleiochasial branches 2–12, each 1–2 times 2-branched; pleiochasial bracts similar in shape and size to distal leaves; dichasial bracts distinct, suborbiculate or reniform, base truncate, margins entire or minutely denticulate, apex obtuse, mucronulate; axillary cymose branches 0–4. |
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Cyathia | peduncle 0.5–1 mm. |
solitary at distal nodes; peduncle 0.3–1(–1.5) mm. |
2n | = 28. |
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Euphorbia myrsinites |
Euphorbia astyla |
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Phenology | Flowering and fruiting spring–summer. | Flowering and fruiting late spring–early fall. |
Habitat | Scrub oak communities, open ground near forests, shrub-steppes. | Desert, grasslands, limestone substrates, usually on very saline or alkaline soils. |
Elevation | 0–2400 m. (0–7900 ft.) | 700–1100 m. (2300–3600 ft.) |
Distribution |
CA; CO; ID; MT; NM; OR; UT; WA; WY; BC; s Europe; w Asia [Introduced in North America]
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TX; Mexico (Coahuila, Nuevo León) |
Discussion | Euphorbia myrsinites is cultivated in much of the flora area, where it can tolerate cold winters. In some areas, it can locally escape from cultivation. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia astyla is a specialist on halophytic, alkaline soils and is known in the flora area only in part of Pecos County. The species is closely related to E. jejuna but differs in its sessile or sub-sessile leaves with a cordate-auriculate base and involucral gland appendages that are not deeply lobed or cleft. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 305. | FNA vol. 12, p. 261. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Tithymalus myrsinites | Chamaesyce astyla |
Name authority | Linnaeus: Sp. Pl. 1: 461. (1753) | Engelmann ex Boissier: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 15(2): 40. (1862) |
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