Euphorbia marginata |
Euphorbia porteriana |
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euphorbe marginée, mountain snow spurge, smoke-on-the-prairie, snow-on-the-mountain, variegated spurge, whitemargined spurge |
Porter's sandmat |
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Habit | Herbs, annual, with taproot. | Herbs, usually annual, sometimes perennial, with slender to occasionally thickened taproot, 5 mm diam.. |
Stems | erect, unbranched or branched, 30–85(–150) cm, pilose or glabrous. |
erect to ascending, sometimes slightly woody at base, not mat-forming, 15–60 cm, usually glabrous, young branches rarely very sparsely short pubescent. |
Leaves | alternate; stipules 0.1–0.3 mm; petiole 0.2–3 mm, glabrous or minutely pilose; blade broadly ovate to elliptic, 32–62(–82) × 18–28(–52) mm, base rounded to cuneate, margins entire, often white on distal leaves, apex acute, rarely mucronate, surfaces glabrous; venation obscure, only midvein conspicuous. |
opposite; stipules usually distinct, occasionally connate basally, triangular, entire or 2–4 parted, apically ciliate, 0.3–0.4 mm, usually glabrous, rarely very sparsely short pubescent; petiole 0.5–1.5 mm, usually glabrous, rarely very sparsely short pubescent; blade ovate, oval, elliptic, oblong-elliptic, or linear-elliptic, 4–12 × 1–7 mm, base asymmetric, rounded or subcordate, margins usually entire, rarely sparsely serrate, apex obtuse to acute, sometimes mucronate, surfaces abaxially often purple or red, adaxially green, usually glabrous, rarely very sparsely short pubescent; 3-veined from base, only midvein conspicuous. |
Involucre | campanulate, 2.2–3.5 × 1.3–2.3 mm, margin between glands deeply divided into fringe of fimbriate lobes, pilose; glands 4–5, green to greenish yellow, reniform to subcircular, 0.7–1.1 × 1–1.6 mm; appendages white, orbiculate, 1.5–2.7 × 1.9–2.9(–3.6) mm, entire. |
turbinate to campanulate, 0.7–1 × 0.8–1 mm, usually glabrous, rarely very sparsely short pubescent; glands 4, brown, slightly concave, elliptic-oblong, 0.1–0.2 × 0.3–0.6 mm; appendages white or pink to dark red, usually oblong or flabellate, occasionally rudimentary and forming narrow rim at edge of gland, (0–)0.1–0.2 × 0.4–1 mm, distal margin entire or crenulate. |
Staminate flowers | 30–70. |
8–14. |
Pistillate flowers | ovary pilose; styles 1–2.5 mm, 2-fid 1/2–2/3 length. |
ovary glabrous, subtended by triangular calyxlike structure; styles 0.4–0.6 mm, 2-fid 1/2 length. |
Capsules | oblate, 3–5 × 3.5–7.5 mm, moderately to densely pilose; columella 3–4.1 mm. |
broadly ovoid, 1–1.5 × 1.7–2.1 mm, glabrous; columella 1–1.4 mm. |
Seeds | orange-tan to gray, ovoid, 3.7–3.9 × 3–3.3 mm, rugose, with 2 transverse ridges (one dark orange to brown, other inconspicuous); caruncle absent. |
reddish brown, ovoid, 4-angled in cross section, 0.7–1 × 0.6–0.7 mm, smooth or obscurely wrinkled. |
Cyathia | in terminal pleiochasia, dichasial bracts narrowly elliptic to oblanceolate with conspicuous white margins; peduncle 1.8–2.7(–22) mm, densely pilose. |
solitary at distal nodes; peduncle 0.5–1.1 mm. |
2n | = 56. |
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Euphorbia marginata |
Euphorbia porteriana |
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Phenology | Flowering and fruiting summer–fall. | Flowering and fruiting year-round. |
Habitat | Disturbed areas and grasslands. | Pine rocklands, coastal scrub, open hammocks. |
Elevation | 0–1700 m. (0–5600 ft.) | 0–10 m. (0–0 ft.) |
Distribution |
AR; CA; CO; CT; IA; IL; IN; KS; LA; MA; MD; MI; MN; MO; MT; NC; NE; NH; NM; NY; OH; OK; PA; RI; SC; SD; TN; TX; UT; VA; WI; WV; WY; MB; ON; QC; SK; c Mexico; s Mexico
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FL |
Discussion | Euphorbia marginata is native to the central United States. The type specimen was collected by Meriwether Lewis along the Yellowstone River in southern Montana in 1806, and it has been reported to be native as far south as Arizona, New Mexico, and Texas, and as far east as southern Minnesota, western Iowa, and Missouri (G. Yatskievych 1999–2013, vol. 2). It is presumably naturalized outside of this area. Euphorbia marginata is widely cultivated as an ornamental for its showy, white-margined distal leaves, and it can escape locally. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia porteriana is found in Miami-Dade County and on Big Pine Key, Monroe County. This species is closely related to E. blodgettii, E. garberi, and E. serpens. The capsules of E. porteriana are almost always completely glabrous, but the authors have seen three specimens with just a few scattered hairs on the keels. A. Herndon (1993) synonymized E. porteriana var. keyensis under E. garberi because of its uniformly hairy capsules, and that treatment is followed here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 248. | FNA vol. 12, p. 285. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Alectoroctonum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | E. bonplandii, Lepadena marginata | Chamaesyce porteriana, C. porteriana var. scoparia, C. scoparia |
Name authority | Pursh: Fl. Amer. Sept. 2: 607. (1813) | (Small) Oudejans: Phytologia 67: 48. (1989) |
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