Euphorbia marginata |
Euphorbia polygonifolia |
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euphorbe marginée, mountain snow spurge, smoke-on-the-prairie, snow-on-the-mountain, variegated spurge, whitemargined spurge |
dune spurge, euphorbe à feuilles de renouée, seaside sandmat, seaside spurge |
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Habit | Herbs, annual, with taproot. | Herbs, annual, with taproot. |
Stems | erect, unbranched or branched, 30–85(–150) cm, pilose or glabrous. |
usually prostrate, occasionally ascending, 5–30 cm, glabrous. |
Leaves | alternate; stipules 0.1–0.3 mm; petiole 0.2–3 mm, glabrous or minutely pilose; blade broadly ovate to elliptic, 32–62(–82) × 18–28(–52) mm, base rounded to cuneate, margins entire, often white on distal leaves, apex acute, rarely mucronate, surfaces glabrous; venation obscure, only midvein conspicuous. |
opposite; stipules usually distinct, occasionally connate basally (distal portion of stem), triangular-subulate, entire or divided, 0.8–1.5 mm, glabrous; petiole 1–3 mm, glabrous; blade oblong, linear-oblong, or linear-lanceolate, 5–16 × 2–4 mm, base slightly asymmetric, obtuse or subcordate, margins entire, apex obtuse, often mucronulate, surfaces uniformly green or reddish tinged, glabrous; venation obscure. |
Involucre | campanulate, 2.2–3.5 × 1.3–2.3 mm, margin between glands deeply divided into fringe of fimbriate lobes, pilose; glands 4–5, green to greenish yellow, reniform to subcircular, 0.7–1.1 × 1–1.6 mm; appendages white, orbiculate, 1.5–2.7 × 1.9–2.9(–3.6) mm, entire. |
obconic-campanulate, 1.2–1.7 × 1–1.4 mm, glabrous; glands 4, occasionally rudimentary, green-yellow to tan or orange-tinged, occasionally shortly stipitate, usually broadly oval to subcircular, sometimes figure eight-shaped, shallowly cupped, 0.1–0.3 × 0.2–0.4 mm; appendages absent or rudimentary. |
Staminate flowers | 30–70. |
5–14. |
Pistillate flowers | ovary pilose; styles 1–2.5 mm, 2-fid 1/2–2/3 length. |
ovary glabrous; styles 0.7–1 mm, 2-fid 1/2 length. |
Capsules | oblate, 3–5 × 3.5–7.5 mm, moderately to densely pilose; columella 3–4.1 mm. |
broadly ovoid, 3–3.5(–4) × (2–)2.4–3 mm, glabrous; columella 2–3 mm. |
Seeds | orange-tan to gray, ovoid, 3.7–3.9 × 3–3.3 mm, rugose, with 2 transverse ridges (one dark orange to brown, other inconspicuous); caruncle absent. |
ashy white, wedge-shaped to slightly ovoid, weakly dorsiventrally compressed and elliptic-terete to bluntly subangled in cross section, back strongly rounded, face slightly rounded, (2–)2.2–2.8 × 1.6–1.9 mm, smooth or minutely pitted, with smooth brown line from top to bottom on adaxial side. |
Cyathia | in terminal pleiochasia, dichasial bracts narrowly elliptic to oblanceolate with conspicuous white margins; peduncle 1.8–2.7(–22) mm, densely pilose. |
solitary or in small, cymose clusters at distal nodes; peduncle 0.5–5 mm. |
2n | = 56. |
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Euphorbia marginata |
Euphorbia polygonifolia |
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Phenology | Flowering and fruiting summer–fall. | Flowering and fruiting early summer–fall. |
Habitat | Disturbed areas and grasslands. | Sandy maritime and freshwater beaches and foredunes. |
Elevation | 0–1700 m. (0–5600 ft.) | 0–200 m. (0–700 ft.) |
Distribution |
AR; CA; CO; CT; IA; IL; IN; KS; LA; MA; MD; MI; MN; MO; MT; NC; NE; NH; NM; NY; OH; OK; PA; RI; SC; SD; TN; TX; UT; VA; WI; WV; WY; MB; ON; QC; SK; c Mexico; s Mexico
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CT; DE; FL; GA; IL; IN; MA; MD; ME; MI; NC; NH; NJ; NY; OH; PA; RI; SC; VA; WI; NB; NS; ON; PE; QC
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Discussion | Euphorbia marginata is native to the central United States. The type specimen was collected by Meriwether Lewis along the Yellowstone River in southern Montana in 1806, and it has been reported to be native as far south as Arizona, New Mexico, and Texas, and as far east as southern Minnesota, western Iowa, and Missouri (G. Yatskievych 1999–2013, vol. 2). It is presumably naturalized outside of this area. Euphorbia marginata is widely cultivated as an ornamental for its showy, white-margined distal leaves, and it can escape locally. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia polygonifolia is native to coastal beaches and dunes along the Atlantic Ocean from the maritime provinces of Canada south to northern Florida. The species also occurs disjunctly along the shores of the North American Great Lakes. Euphorbia polygonifolia was native to Quebec but is now considered extirpated from the province. It has apparently been introduced in Europe, but it is unclear if it has persisted there (L. C. Wheeler 1941). Immature individuals of this species can be somewhat difficult to distinguish from E. bombensis where their ranges overlap (Virginia to northern Florida). Where they occur together, E. polygonifolia tends to be a pioneer species on the upper beach and foredune front, whereas E. bombensis tends to inhabit areas behind the foredune (R. D. Porcher and D. A. Rayner 2002). Euphorbia polygonifolia can be distinguished also by its larger capsules and larger, wedge-shaped to slightly ovoid seeds. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 248. | FNA vol. 12, p. 284. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Alectoroctonum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | E. bonplandii, Lepadena marginata | Chamaesyce polygonifolia |
Name authority | Pursh: Fl. Amer. Sept. 2: 607. (1813) | Linnaeus: Sp. Pl. 1: 455. (1753) |
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