Euphorbia maculata |
Euphorbia yaquiana |
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euphorbe maculée, milk purslane, milk spurge, prostrate spurge, sandmat, spotted sandmat, spotted sandmat or spurge, spotted spurge |
hairy Mojave spurge |
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Habit | Herbs, annual, with taproot. | Herbs, perennial, with thick rootstock. |
Stems | usually prostrate, occasionally with ascending tips, often mat-forming, not rooting at nodes, 5–45 cm, densely and evenly short-sericeous to sericeous or villous. |
slender, erect or ascending, sometimes sinuous, densely branched near base, 10–50 cm, moderately to densely puberulent to lanulose. |
Leaves | opposite; stipules distinct, linear-subulate, sometimes irregularly 2–3-lobed, 1–1.3 mm, sparsely short-sericeous to sericeous or villous; petiole 0.5–1.5 mm, moderately short-sericeous to sericeous or villous; blade oblong-ovate to ovate-elliptic or oblong-elliptic, 4–18 × 2.5–8 mm, base strongly asymmetric, one side usually angled and other ± truncate and expanded into small, rounded auricle, margins serrulate (longer side) or subentire (shorter side), apex rounded or broadly acute, abaxial surface pale grayish green, moderately to densely lanulose to villous, adaxial surface usually with irregular reddish streak along midvein, glabrate or with sparse, long, slender hairs; palmately veined at base, pinnate distally. |
petiole 0–1 mm; blade usually lanceolate or elliptic-lanceolate, sometimes slightly oblanceolate, 8–30 × 6–14 mm, base usually acute, occasionally short-attenuate, rarely obtuse, margins entire, apex usually acute, occasionally obtuse, acuminate, or cuspidate, surfaces sparsely to moderately puberulent to lanulose; venation pinnate, sometimes obscure, midvein prominent. |
Involucre | obconic, 0.8–1 × 0.6–0.8 mm, sparsely strigose to short-sericeous; glands 4, green to yellow-green, turning pink with age, usually ± unequal, narrowly oblong to nearly linear, 0.1–0.2 × 0.2–0.5 mm; appendages white to reddish tinged, lunate to oblong, 0.1–0.3 × 0.2–1.5 mm, distal margin crenulate. |
campanulate to broadly turbinate, 2.2–3 × 2–2.5 mm, puberulent to lanulose; glands 4, semicircular, trapezoidal, or elliptic-truncate, 0.8–1.5 × 1–2.2 mm, margins strongly crenate or dentate; horns usually absent, if present then straight, 0.1–0.2 mm, generally equaling teeth on gland margin. |
Staminate flowers | 2–5. |
12–20. |
Pistillate flowers | ovary sericeous; styles 0.3–0.4 mm, 2-fid at apex. |
ovary usually puberulent, occasionally lanulose; styles 1–1.2 mm, 2-fid. |
Capsules | ovoid, well exserted from involucre at maturity, 1.3–1.5 × 1.2–1.4 mm, sparsely to moderately and evenly sericeous; columella 1–1.2 mm. |
oblong-ovoid, 3.5–4 × 3–4 mm, 3-lobed; cocci rounded, smooth, usually puberulent, occasionally lanulose; columella 2.5–3 mm. |
Seeds | white to light brown, oblong-ovoid, sharply angular in cross section, 1–1.2 × 0.6–0.9 mm, with 3–4 low, transverse ridges that cross angles. |
gray to whitish, oblong cylindric, 2–3 × 1.5–1.8 mm, irregularly shallowly pitted to almost smooth; caruncle conic, 0.6 × 0.6 mm. |
Cyathia | solitary or in small, cymose clusters at distal nodes or on congested, axillary branches; peduncle 0.1–0.6 mm. |
peduncle 0.3–0.8 mm. |
Cyathial | arrangement: terminal pleiochasial branches 3–5, each 1–2 times 2-branched; pleiochasial bracts broadly ovate to subcordate, usually similar in size to, occasionally wider than, distal leaves; dichasial bracts distinct, broadly ovate to almost reniform, base obtuse, margins entire, apex obtuse, acuminate to cuspidate; axillary cymose branches 0–5. |
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2n | = 28. |
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Euphorbia maculata |
Euphorbia yaquiana |
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Phenology | Flowering and fruiting spring–fall. | Flowering and fruiting spring–summer. |
Habitat | Disturbed areas, fallow fields, gardens, sidewalk cracks, railroads, roadsides. | Ponderosa pine forests, oak-pine mixed forests, dry stream banks and beds, open scrub areas, roadsides. |
Elevation | 0–1500 m. (0–4900 ft.) | 1000–2200 m. (3300–7200 ft.) |
Distribution |
AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; BC; NB; NS; ON; PE; QC; Mexico; Central America; South America; West Indies; Eurasia; Africa; Pacific Islands (New Zealand); Australia
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AZ |
Discussion | Euphorbia maculata is a widespread weed in temperate latitudes, and it also occurs in cool climates at higher elevations in the tropics. It is presumed to be native to eastern and central North America, but given its extremely weedy tendencies, it is difficult to know for sure. It spreads readily in association with greenhouse plants and earth-moving activities, and it is notorious for its ability to colonize sidewalk cracks in the summer, even in congested cities. The name E. maculata was misapplied by most earlier botanists (for example, L. C. Wheeler 1941) to plants with ascending stems that are treated here as E. nutans. D. G. Burch (1966) reviewed the sources of data used by Linnaeus in his original description and concluded that the name E. maculata applies to this prostrate-stemmed taxon. For further discussion of the distinctions between E. maculata and the similar 47. E. humistrata, see the treatment of that species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia yaquiana is endemic to Pima and Graham counties in southern Arizona and is known only from the Santa Catalina and Pinaleño mountains. Records of E. yaquiana from southwestern Colorado (as E. incisa var. mollis) likely represent misidentifications of E. brachycera; therefore, those disjunct occurrences have been excluded here from the distribution of E. yaquiana. Euphorbia yaquiana has often been treated as a synonym of E. schizoloba var. mollis, but molecular phylogenetic data show that it is more closely related to E. brachycera and E. chamaesula (J. A. Peirson et al. 2014). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 275. | FNA vol. 12, p. 313. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > subg. Esula |
Sibling taxa | ||
Synonyms | Chamaesyce maculata, C. mathewsii, C. supina, C. tracyi, E. supina | E. schizoloba var. mollis, E. incisa var. mollis |
Name authority | Linnaeus: Sp. Pl. 1: 455. (1753) | Tidestrom: Proc. Biol. Soc. Wash. 48: 41. (1935) |
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