Euphorbia maculata |
Euphorbia parishii |
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euphorbe maculée, milk purslane, milk spurge, prostrate spurge, sandmat, spotted sandmat, spotted sandmat or spurge, spotted spurge |
Parish's sandmat |
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Habit | Herbs, annual, with taproot. | Herbs or subshrubs, perennial, with thickened and often woody rootstock. |
Stems | usually prostrate, occasionally with ascending tips, often mat-forming, not rooting at nodes, 5–45 cm, densely and evenly short-sericeous to sericeous or villous. |
prostrate, sometimes forming dense mounds, 10–50 cm, glabrous. |
Leaves | opposite; stipules distinct, linear-subulate, sometimes irregularly 2–3-lobed, 1–1.3 mm, sparsely short-sericeous to sericeous or villous; petiole 0.5–1.5 mm, moderately short-sericeous to sericeous or villous; blade oblong-ovate to ovate-elliptic or oblong-elliptic, 4–18 × 2.5–8 mm, base strongly asymmetric, one side usually angled and other ± truncate and expanded into small, rounded auricle, margins serrulate (longer side) or subentire (shorter side), apex rounded or broadly acute, abaxial surface pale grayish green, moderately to densely lanulose to villous, adaxial surface usually with irregular reddish streak along midvein, glabrate or with sparse, long, slender hairs; palmately veined at base, pinnate distally. |
opposite; stipules distinct, subulate-filiform, 0.3–0.9 mm, pilose; petiole 0.3–1.2 mm, glabrous; blade usually ovate, rarely oblong, 2–7 × 1–5 mm, base usually asymmetric, rounded to hemicordate, margins entire, apex usually obtuse, rarely acute, surfaces glabrous; only midvein conspicuous. |
Involucre | obconic, 0.8–1 × 0.6–0.8 mm, sparsely strigose to short-sericeous; glands 4, green to yellow-green, turning pink with age, usually ± unequal, narrowly oblong to nearly linear, 0.1–0.2 × 0.2–0.5 mm; appendages white to reddish tinged, lunate to oblong, 0.1–0.3 × 0.2–1.5 mm, distal margin crenulate. |
obconic to campanulate, 1–1.4 × 0.9–1.3 mm, glabrous except for pilose lobes; glands 4, pink to maroon, circular, 0.3–0.4 × 0.3–0.4 mm; appendages absent. |
Staminate flowers | 2–5. |
40–50. |
Pistillate flowers | ovary sericeous; styles 0.3–0.4 mm, 2-fid at apex. |
ovary glabrous; styles 0.3–0.6 mm, 2-fid 1/2 length. |
Capsules | ovoid, well exserted from involucre at maturity, 1.3–1.5 × 1.2–1.4 mm, sparsely to moderately and evenly sericeous; columella 1–1.2 mm. |
ovoid to oblate-ovoid, 1.6–1.7 × 1.6–1.9 mm, glabrous; columella 1.2–1.5 mm. |
Seeds | white to light brown, oblong-ovoid, sharply angular in cross section, 1–1.2 × 0.6–0.9 mm, with 3–4 low, transverse ridges that cross angles. |
whitish to light brown, ovoid, 4-angled in cross section, 1.2–1.4 × 0.6–0.8 mm, rugose or with indistinct, irregular, low transverse ridges. |
Cyathia | solitary or in small, cymose clusters at distal nodes or on congested, axillary branches; peduncle 0.1–0.6 mm. |
solitary at distal nodes; peduncle 0.1–0.6(–2.2) mm. |
2n | = 28. |
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Euphorbia maculata |
Euphorbia parishii |
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Phenology | Flowering and fruiting spring–fall. | Flowering and fruiting fall–summer. |
Habitat | Disturbed areas, fallow fields, gardens, sidewalk cracks, railroads, roadsides. | Desert scrub, often with creosote bush, disturbed roadsides, rocky soils. |
Elevation | 0–1500 m. (0–4900 ft.) | -90–600 m. (-300–2000 ft.) |
Distribution |
AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; BC; NB; NS; ON; PE; QC; Mexico; Central America; South America; West Indies; Eurasia; Africa; Pacific Islands (New Zealand); Australia
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CA; NV |
Discussion | Euphorbia maculata is a widespread weed in temperate latitudes, and it also occurs in cool climates at higher elevations in the tropics. It is presumed to be native to eastern and central North America, but given its extremely weedy tendencies, it is difficult to know for sure. It spreads readily in association with greenhouse plants and earth-moving activities, and it is notorious for its ability to colonize sidewalk cracks in the summer, even in congested cities. The name E. maculata was misapplied by most earlier botanists (for example, L. C. Wheeler 1941) to plants with ascending stems that are treated here as E. nutans. D. G. Burch (1966) reviewed the sources of data used by Linnaeus in his original description and concluded that the name E. maculata applies to this prostrate-stemmed taxon. For further discussion of the distinctions between E. maculata and the similar 47. E. humistrata, see the treatment of that species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia parishii is common in the Death Valley region of southern California, where it is often encountered well below sea level. The species is frequently confused with E. micromera and E. polycarpa but differs from the former in being a more robust plant with larger cyathia and from the latter in lacking involucral gland appendages. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 275. | FNA vol. 12, p. 280. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Chamaesyce maculata, C. mathewsii, C. supina, C. tracyi, E. supina | Chamaesyce parishii, E. polycarpa var. parishii |
Name authority | Linnaeus: Sp. Pl. 1: 455. (1753) | Greene: Bull. Calif. Acad. Sci. 2: 56. (1886) |
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