Euphorbia maculata |
Euphorbia innocua |
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euphorbe maculée, milk purslane, milk spurge, prostrate spurge, sandmat, spotted sandmat, spotted sandmat or spurge, spotted spurge |
velvet spurge |
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Habit | Herbs, annual, with taproot. | Herbs, perennial, with moderately to strongly thickened rootstock. |
Stems | usually prostrate, occasionally with ascending tips, often mat-forming, not rooting at nodes, 5–45 cm, densely and evenly short-sericeous to sericeous or villous. |
prostrate to decumbent or ascending, branched (often near base), 7–45 cm, densely pilose. |
Leaves | opposite; stipules distinct, linear-subulate, sometimes irregularly 2–3-lobed, 1–1.3 mm, sparsely short-sericeous to sericeous or villous; petiole 0.5–1.5 mm, moderately short-sericeous to sericeous or villous; blade oblong-ovate to ovate-elliptic or oblong-elliptic, 4–18 × 2.5–8 mm, base strongly asymmetric, one side usually angled and other ± truncate and expanded into small, rounded auricle, margins serrulate (longer side) or subentire (shorter side), apex rounded or broadly acute, abaxial surface pale grayish green, moderately to densely lanulose to villous, adaxial surface usually with irregular reddish streak along midvein, glabrate or with sparse, long, slender hairs; palmately veined at base, pinnate distally. |
alternate; stipules to 0.1 mm; petiole (0.7–)1.1–3.5 mm, pilose; blade ovate to orbiculate, 4.6–17(–25) × 4.5–15(–19) mm, base cordate, margins entire, apex rounded to obtuse, surfaces densely pilose; venation obscure, usually only midvein conspicuous. |
Involucre | obconic, 0.8–1 × 0.6–0.8 mm, sparsely strigose to short-sericeous; glands 4, green to yellow-green, turning pink with age, usually ± unequal, narrowly oblong to nearly linear, 0.1–0.2 × 0.2–0.5 mm; appendages white to reddish tinged, lunate to oblong, 0.1–0.3 × 0.2–1.5 mm, distal margin crenulate. |
campanulate, 1–1.3 × 1.2–1.4 mm, pilose; glands 4, yellow to green, elliptic, 0.2–0.3 × 0.5–0.6 mm; appendages green, elliptic, 0.4–0.5 × 0.5–0.9 mm, entire or crenulate, ciliate. |
Staminate flowers | 2–5. |
5–10. |
Pistillate flowers | ovary sericeous; styles 0.3–0.4 mm, 2-fid at apex. |
ovary pilose; styles 0.4–0.7 mm, 2-fid 1/2 length. |
Capsules | ovoid, well exserted from involucre at maturity, 1.3–1.5 × 1.2–1.4 mm, sparsely to moderately and evenly sericeous; columella 1–1.2 mm. |
depressed-ovoid, 2–2.5 × 2.7–3.3 mm, pilose; columella 1.6–2.1 mm. |
Seeds | white to light brown, oblong-ovoid, sharply angular in cross section, 1–1.2 × 0.6–0.9 mm, with 3–4 low, transverse ridges that cross angles. |
gray to brown, ovoid, 1.5–1.7 × 1.2–1.3 mm, rugose with whitish ridges; caruncle absent. |
Cyathia | solitary or in small, cymose clusters at distal nodes or on congested, axillary branches; peduncle 0.1–0.6 mm. |
in terminal dichasia (often weakly defined); peduncle 1–2.7 mm, densely pilose. |
2n | = 28. |
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Euphorbia maculata |
Euphorbia innocua |
|
Phenology | Flowering and fruiting spring–fall. | Flowering and fruiting early winter–late spring. |
Habitat | Disturbed areas, fallow fields, gardens, sidewalk cracks, railroads, roadsides. | Sandy soils or dunes, grasslands, pastures. |
Elevation | 0–1500 m. (0–4900 ft.) | 0–20 m. (0–100 ft.) |
Distribution |
AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; BC; NB; NS; ON; PE; QC; Mexico; Central America; South America; West Indies; Eurasia; Africa; Pacific Islands (New Zealand); Australia
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TX |
Discussion | Euphorbia maculata is a widespread weed in temperate latitudes, and it also occurs in cool climates at higher elevations in the tropics. It is presumed to be native to eastern and central North America, but given its extremely weedy tendencies, it is difficult to know for sure. It spreads readily in association with greenhouse plants and earth-moving activities, and it is notorious for its ability to colonize sidewalk cracks in the summer, even in congested cities. The name E. maculata was misapplied by most earlier botanists (for example, L. C. Wheeler 1941) to plants with ascending stems that are treated here as E. nutans. D. G. Burch (1966) reviewed the sources of data used by Linnaeus in his original description and concluded that the name E. maculata applies to this prostrate-stemmed taxon. For further discussion of the distinctions between E. maculata and the similar 47. E. humistrata, see the treatment of that species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia innocua is restricted to south coastal Texas in Aransas, Calhoun, Kenedy, Kleberg, Nueces, Refugio, San Patricio, and Willacy counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 275. | FNA vol. 12, p. 247. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Alectoroctonum |
Sibling taxa | ||
Synonyms | Chamaesyce maculata, C. mathewsii, C. supina, C. tracyi, E. supina | |
Name authority | Linnaeus: Sp. Pl. 1: 455. (1753) | L. C. Wheeler: Contr. Gray Herb. 127: 62, plate 3, fig. D. (1939) |
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