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euphorbe maculée, milk purslane, milk spurge, prostrate spurge, sandmat, spotted sandmat, spotted sandmat or spurge, spotted spurge

alkali spurge, Pecos spurge

Habit Herbs, annual, with taproot. Herbs, perennial, with woody or fibrous-fleshy taproot, 5–12 mm thick.
Stems

usually prostrate, occasionally with ascending tips, often mat-forming, not rooting at nodes, 5–45 cm, densely and evenly short-sericeous to sericeous or villous.

decumbent, ascending, or erect, few to many emerging from woody crown, 5–25(–50) cm, glabrous.

Leaves

opposite;

stipules distinct, linear-subulate, sometimes irregularly 2–3-lobed, 1–1.3 mm, sparsely short-sericeous to sericeous or villous;

petiole 0.5–1.5 mm, moderately short-sericeous to sericeous or villous;

blade oblong-ovate to ovate-elliptic or oblong-elliptic, 4–18 × 2.5–8 mm, base strongly asymmetric, one side usually angled and other ± truncate and expanded into small, rounded auricle, margins serrulate (longer side) or subentire (shorter side), apex rounded or broadly acute, abaxial surface pale grayish green, moderately to densely lanulose to villous, adaxial surface usually with irregular reddish streak along midvein, glabrate or with sparse, long, slender hairs; palmately veined at base, pinnate distally.

opposite;

stipules connate into deltate scale, 0.2–0.5 mm, minutely lacerate at apex, glabrous;

petiole 0–0.2(–0.3) mm, glabrous;

blade orbiculate-reniform to acute-cordate, 2–5(–8) × 2–5(–6) mm, base ± asymmetric, cordate to auriculate, sometimes clasping stem, margins entire, apex narrowly acute, surfaces glabrous; 2- or 3-veined from base, but usually only midvein conspicuous.

Involucre

obconic, 0.8–1 × 0.6–0.8 mm, sparsely strigose to short-sericeous;

glands 4, green to yellow-green, turning pink with age, usually ± unequal, narrowly oblong to nearly linear, 0.1–0.2 × 0.2–0.5 mm;

appendages white to reddish tinged, lunate to oblong, 0.1–0.3 × 0.2–1.5 mm, distal margin crenulate.

broadly campanulate, 0.8–1.4 × 0.9–1.4 mm, glabrous;

glands 4, yellow-green to brownish, oblong, 0.2–0.3 × 0.5–0.7 mm;

appendages white, flabellate to oblong, 0.1–0.2(–0.5) × 0.4–0.8 mm, distal margin entire or dentate-crenate.

Staminate flowers

2–5.

22–26.

Pistillate flowers

ovary sericeous;

styles 0.3–0.4 mm, 2-fid at apex.

ovary glabrous;

styles 0.3–0.4 mm, unbranched, thickened-clavate.

Capsules

ovoid, well exserted from involucre at maturity, 1.3–1.5 × 1.2–1.4 mm, sparsely to moderately and evenly sericeous;

columella 1–1.2 mm.

ovoid and broadly triangular, 1.5–1.9(–2.5) × 1.4–1.6(–2.2) mm, glabrous;

columella 1.2–1.8 mm.

Seeds

white to light brown, oblong-ovoid, sharply angular in cross section, 1–1.2 × 0.6–0.9 mm, with 3–4 low, transverse ridges that cross angles.

white, oblong, 4-angled in cross section, adaxial faces slightly concave, with long raphe between, 1.5–1.8 × 0.7–1 mm, markedly foveolate, with irregular to ± parallel or anastomosing ridges.

Cyathia

solitary or in small, cymose clusters at distal nodes or on congested, axillary branches;

peduncle 0.1–0.6 mm.

solitary at distal nodes;

peduncle 0.3–1(–1.5) mm.

2n

= 28.

= 28.

Euphorbia maculata

Euphorbia astyla

Phenology Flowering and fruiting spring–fall. Flowering and fruiting late spring–early fall.
Habitat Disturbed areas, fallow fields, gardens, sidewalk cracks, railroads, roadsides. Desert, grasslands, limestone substrates, usually on very saline or alkaline soils.
Elevation 0–1500 m. (0–4900 ft.) 700–1100 m. (2300–3600 ft.)
Distribution
from FNA
AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; BC; NB; NS; ON; PE; QC; Mexico; Central America; South America; West Indies; Eurasia; Africa; Pacific Islands (New Zealand); Australia
[WildflowerSearch map]
[BONAP county map]
from FNA
TX; Mexico (Coahuila, Nuevo León)
[BONAP county map]
Discussion

Euphorbia maculata is a widespread weed in temperate latitudes, and it also occurs in cool climates at higher elevations in the tropics. It is presumed to be native to eastern and central North America, but given its extremely weedy tendencies, it is difficult to know for sure. It spreads readily in association with greenhouse plants and earth-moving activities, and it is notorious for its ability to colonize sidewalk cracks in the summer, even in congested cities. The name E. maculata was misapplied by most earlier botanists (for example, L. C. Wheeler 1941) to plants with ascending stems that are treated here as E. nutans. D. G. Burch (1966) reviewed the sources of data used by Linnaeus in his original description and concluded that the name E. maculata applies to this prostrate-stemmed taxon. For further discussion of the distinctions between E. maculata and the similar 47. E. humistrata, see the treatment of that species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia astyla is a specialist on halophytic, alkaline soils and is known in the flora area only in part of Pecos County. The species is closely related to E. jejuna but differs in its sessile or sub-sessile leaves with a cordate-auriculate base and involucral gland appendages that are not deeply lobed or cleft.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 275. FNA vol. 12, p. 261.
Parent taxa Euphorbiaceae > Euphorbia > sect. Anisophyllum Euphorbiaceae > Euphorbia > sect. Anisophyllum
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms Chamaesyce maculata, C. mathewsii, C. supina, C. tracyi, E. supina Chamaesyce astyla
Name authority Linnaeus: Sp. Pl. 1: 455. (1753) Engelmann ex Boissier: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 15(2): 40. (1862)
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