Euphorbia lurida |
Euphorbia stictospora |
|
---|---|---|
woodland spurge |
mat spurge, narrow-seed spurge, slimseed sandmat |
|
Habit | Herbs, perennial, with thick rootstock. | Herbs, annual, with taproot. |
Stems | erect or ascending, unbranched, sometimes sinuous, 5–30 cm, glabrous or sparsely to densely puberulent. |
prostrate, often mat-forming, occasionally with ascending tips, 5–45 cm, densely and evenly pilose to lanate. |
Leaves | petiole 0–1 mm; blade oblanceolate to obovate, 8–20 mm × 3–7 mm, base truncate or cuneate, margins entire, apex obtuse to rounded, minutely mucronate, surfaces puberulent or glabrous; venation inconspicuous, only midvein prominent. |
opposite; stipules distinct or connate basally on one side of stem, entire or irregularly toothed or fringed, 0.5–1.2 mm, pilose to lanate; petiole 0.3–1.5 mm, pilose to lanate; blade usually oblong to oblong-obovate, occasionally nearly circular, 3–10(–15) × 2–5(–10) mm, base asymmetric, one side usually angled or rounded and other truncate-auriculate, margins minutely or conspicuously serrulate at least toward apex, apex usually broadly rounded to broadly acute, occasionally emarginate, abaxial surface often ± lighter green and without reddish spot, both surfaces sparsely to moderately pilose to lanate; 3-veined from base or venation obscure. |
Involucre | cupulate, 2–2.2 × 1.3–1.8 mm, glabrous; glands 4, oblong to broadly ovate, usually truncate, 0.5–0.8 × 1–1.6 mm, margins irregularly crenate to strongly dentate; horns absent or usually divergent or straight, 0.1–0.3 mm, usually slightly longer than, occasionally equaling, teeth on gland margin. |
obconic, 0.7–1 × 0.4–0.6 mm, moderately to densely pilose to lanate; glands 4, reddish, ± unequal, oblong, 0.1 × 0.1–0.3 mm; appendages white to strongly pinkish or reddish tinged, often unequal, sometimes 1 to all absent, 0–0.3 × 0–0.4 mm, 3-lobed or rudimentarily 1-lobed, distal margin crenate. |
Staminate flowers | 10–20. |
3–9. |
Pistillate flowers | ovary glabrous or puberulent; styles 0.7–1 mm, 2-fid. |
ovary pilose to villous, hairs occasionally slightly appressed; styles 0.2–0.5 mm, unbranched or inconspicuously notched at tip. |
Capsules | ovoid, 3.5–4 × 4–4.5 mm, 3-lobed; cocci rounded, smooth to slightly rugose, glabrous; columella 3.2–3.5 mm. |
ovoid, 1.6–2.3 × 1.4–1.5 mm, moderately to densely villous with hairs usually slightly appressed, pubescence often concentrated on proximal 1/2 or along lobes; columella 1.5–2 mm. |
Seeds | gray to dark gray, truncate-oblong to truncate-ovoid, 2.8–3 × 1.7–2 mm, irregularly pitted; caruncle conic, 0.6 × 0.7 mm. |
light to dark brown, usually mottled, sometimes with thin, white coating, often wearing away irregularly, narrowly oblong-ovoid to ellipsoid, 3–4-angled in cross section, 1–1.5 × 0.5–0.6 mm, with short, irregularly interrupted furrows, appearing partially and irregularly few-ridged. |
Cyathial | arrangement: terminal pleiochasial branches 3–5, each 1–4 times 2-branched; pleiochasial bracts ovate to broadly ovate or oblanceolate, wider than distal leaves; dichasial bracts distinct, rounded, oblanceolate, or subreniform, base cuneate or obtuse, margins entire, apex obtuse, slightly mucronate; axillary cymose branches 0–4. |
|
Cyathia | peduncle 0.3–0.9 mm. |
solitary at leaf nodes or in small, cymose clusters on congested, axillary branches; peduncle 0.7–2.5 mm. |
Euphorbia lurida |
Euphorbia stictospora |
|
Phenology | Flowering and fruiting spring–summer. | Flowering and fruiting midsummer–early fall. |
Habitat | Open pine-oak forests, dry slopes and canyons. | Open disturbed areas, rocky slopes. |
Elevation | 1300–2800 m. (4300–9200 ft.) | 100–2100 m. (300–6900 ft.) |
Distribution |
AZ; CA; NM; NV; UT; Mexico (Baja California)
|
AZ; CO; IA; KS; MO; ND; NE; NM; OK; SD; TX; WY; Mexico (Chihuahua, Durango, San Luis Potosí)
|
Discussion | Euphorbia lurida has been treated as a complex of several taxa in the past, but only a single, broadly-defined species is recognized here. This species is variable in both the pubescence and shape of the bracts subtending the cyathia and also in the degree of crenation of the gland margin. In the northern part of its range, E. lurida appears to intergrade with E. brachycera, and it can be difficult to distinguish these two species in northern Arizona and New Mexico. A report of the species from Sonora, Mexico, based on a single immature collection (V. W. Steinmann and R. S. Felger 1997) has not been verified. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia stictospora has been recorded from New York, but this disjunct occurrence likely represents a waif or misidentification. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 305. | FNA vol. 12, p. 289. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | E. palmeri, E. palmeri var. subpubens, E. subpubens, Tithymalus luridus, T. palmeri, T. subpubens | Chamaesyce stictospora |
Name authority | Engelmann: in J. C. Ives, Rep. Colorado R. 4: 26. (1861) | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 187. (1859) |
Web links |