Euphorbia lurida |
Euphorbia micromera |
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woodland spurge |
desert spurge, Sonoran sandmat, tiny sandmat |
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Habit | Herbs, perennial, with thick rootstock. | Herbs, annual, with slender taproot. |
Stems | erect or ascending, unbranched, sometimes sinuous, 5–30 cm, glabrous or sparsely to densely puberulent. |
prostrate, mat-forming, 5–35 cm, glabrous or shortly pilose. |
Leaves | petiole 0–1 mm; blade oblanceolate to obovate, 8–20 mm × 3–7 mm, base truncate or cuneate, margins entire, apex obtuse to rounded, minutely mucronate, surfaces puberulent or glabrous; venation inconspicuous, only midvein prominent. |
opposite; stipules distinct, subulate, 0.2–0.4 mm, pilose; petiole 0.5–1.2 mm, glabrous or pilose; blade ovate to elliptic, 6–15 × 2–4 mm, base asymmetric, one side cuneate to rounded, other side rounded, margins entire, apex obtuse, surfaces glabrous or pilose; venation obscure or only midvein conspicuous. |
Involucre | cupulate, 2–2.2 × 1.3–1.8 mm, glabrous; glands 4, oblong to broadly ovate, usually truncate, 0.5–0.8 × 1–1.6 mm, margins irregularly crenate to strongly dentate; horns absent or usually divergent or straight, 0.1–0.3 mm, usually slightly longer than, occasionally equaling, teeth on gland margin. |
campanulate, 0.4–0.6 × 0.5–0.9 mm, glabrous or pilose; glands 4, red, circular to oblong, 0.1 × 0.1–0.2 mm; appendages absent. |
Staminate flowers | 10–20. |
2–5. |
Pistillate flowers | ovary glabrous or puberulent; styles 0.7–1 mm, 2-fid. |
ovary usually glabrous, rarely pilose; styles 0.1–0.2 mm, 2-fid at apex. |
Capsules | ovoid, 3.5–4 × 4–4.5 mm, 3-lobed; cocci rounded, smooth to slightly rugose, glabrous; columella 3.2–3.5 mm. |
oblong, 1.3–1.5 × 1.1–1.3 mm, usually glabrous, rarely pilose; columella 1–1.2 mm. |
Seeds | gray to dark gray, truncate-oblong to truncate-ovoid, 2.8–3 × 1.7–2 mm, irregularly pitted; caruncle conic, 0.6 × 0.7 mm. |
light gray, narrowly ovoid, 4-angled in cross section, 0.9–1 × 0.5–0.6 mm, smooth to slightly rugose or with 1–4 faint transverse ridges that do not pass through abaxial keel. |
Cyathial | arrangement: terminal pleiochasial branches 3–5, each 1–4 times 2-branched; pleiochasial bracts ovate to broadly ovate or oblanceolate, wider than distal leaves; dichasial bracts distinct, rounded, oblanceolate, or subreniform, base cuneate or obtuse, margins entire, apex obtuse, slightly mucronate; axillary cymose branches 0–4. |
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Cyathia | peduncle 0.3–0.9 mm. |
solitary at distal nodes; peduncle 0.4–1.4 mm. |
Euphorbia lurida |
Euphorbia micromera |
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Phenology | Flowering and fruiting spring–summer. | Flowering nearly year-round in response to sufficient rainfall. |
Habitat | Open pine-oak forests, dry slopes and canyons. | Desert scrub, riparian woods with ash and willow, saltbush scrub, Joshua tree woodlands and grasslands, often in sandy or gravelly areas. |
Elevation | 1300–2800 m. (4300–9200 ft.) | -20–1800 m. (-100–5900 ft.) |
Distribution |
AZ; CA; NM; NV; UT; Mexico (Baja California)
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AZ; CA; NM; NV; TX; UT; Mexico (Baja California, Baja California Sur, Chihuahua, Coahuila, Durango, Sonora)
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Discussion | Euphorbia lurida has been treated as a complex of several taxa in the past, but only a single, broadly-defined species is recognized here. This species is variable in both the pubescence and shape of the bracts subtending the cyathia and also in the degree of crenation of the gland margin. In the northern part of its range, E. lurida appears to intergrade with E. brachycera, and it can be difficult to distinguish these two species in northern Arizona and New Mexico. A report of the species from Sonora, Mexico, based on a single immature collection (V. W. Steinmann and R. S. Felger 1997) has not been verified. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 12, p. 305. | FNA vol. 12, p. 278. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | E. palmeri, E. palmeri var. subpubens, E. subpubens, Tithymalus luridus, T. palmeri, T. subpubens | Chamaesyce micromera |
Name authority | Engelmann: in J. C. Ives, Rep. Colorado R. 4: 26. (1861) | Boissier: Proc. Amer. Acad. Arts 5: 171. (1861) |
Web links |