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woodland spurge

dwarf broomspurge

Habit Herbs, perennial, with thick rootstock. Herbs, perennial, with woody or fibrous-fleshy, napiform, branched or tuberous taproot, 3–15 mm thick.
Stems

erect or ascending, unbranched, sometimes sinuous, 5–30 cm, glabrous or sparsely to densely puberulent.

ascending to erect, densely emerging from woody crown, 5–15 cm, glabrous.

Leaves

petiole 0–1 mm;

blade oblanceolate to obovate, 8–20 mm × 3–7 mm, base truncate or cuneate, margins entire, apex obtuse to rounded, minutely mucronate, surfaces puberulent or glabrous;

venation inconspicuous, only midvein prominent.

opposite;

stipules connate into lacerate or 2-fid, lanceolate or deltate scale, 0.5–0.9 mm, glabrous;

petiole 0.3–0.9 mm, glabrous;

blade orbiculate-obovate, ovate, or elliptic, 3–6(–8) × 1.8–5 mm, base moderately asymmetric, rounded to truncate, margins entire, apex blunt to acute, surfaces glabrous; 2–3-veined from base but usually only midvein conspicuous.

Involucre

cupulate, 2–2.2 × 1.3–1.8 mm, glabrous;

glands 4, oblong to broadly ovate, usually truncate, 0.5–0.8 × 1–1.6 mm, margins irregularly crenate to strongly dentate;

horns absent or usually divergent or straight, 0.1–0.3 mm, usually slightly longer than, occasionally equaling, teeth on gland margin.

broadly campanulate, 1.2–1.5 × 0.8–1.2 mm, glabrous;

glands 4, yellowish to green or purplish, oblong, 0.2–0.4 × 0.5–1 mm;

appendages erect or spreading, white, 0.3–0.6 × 0.8–1.2 mm, usually deeply dissected into 4–5 acuminate lobes, rarely undivided, when divided distal margin rarely entire or crenate.

Staminate flowers

10–20.

12–35.

Pistillate flowers

ovary glabrous or puberulent;

styles 0.7–1 mm, 2-fid.

ovary glabrous;

styles 0.4–0.5 mm, unbranched thickened-clavate.

Capsules

ovoid, 3.5–4 × 4–4.5 mm, 3-lobed;

cocci rounded, smooth to slightly rugose, glabrous;

columella 3.2–3.5 mm.

ovoid and broadly triangular, 1.8–2.2(–2.7) × 1.5–2.1 mm, glabrous;

columella 1.5–2.2 mm.

Seeds

gray to dark gray, truncate-oblong to truncate-ovoid, 2.8–3 × 1.7–2 mm, irregularly pitted;

caruncle conic, 0.6 × 0.7 mm.

whitish, oblong, 4-angled in cross section, adaxial faces concave, with long raphe between, 1.5–2(–2.3) × 0.6–0.8 mm, dimpled with faint irregular transverse wrinkles or with up to 10 low, rounded transverse ridges.

Cyathial

arrangement: terminal pleiochasial branches 3–5, each 1–4 times 2-branched;

pleiochasial bracts ovate to broadly ovate or oblanceolate, wider than distal leaves;

dichasial bracts distinct, rounded, oblanceolate, or subreniform, base cuneate or obtuse, margins entire, apex obtuse, slightly mucronate;

axillary cymose branches 0–4.

Cyathia

peduncle 0.3–0.9 mm.

solitary at distal nodes;

peduncle 0.8–1.5 mm.

Euphorbia lurida

Euphorbia jejuna

Phenology Flowering and fruiting spring–summer. Flowering and fruiting early spring–summer.
Habitat Open pine-oak forests, dry slopes and canyons. Thin calcareous soils (caliche) on limestone hills.
Elevation 1300–2800 m. (4300–9200 ft.) 500–900 m. (1600–3000 ft.)
Distribution
from FNA
AZ; CA; NM; NV; UT; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
from FNA
TX; Mexico (Chihuahua, Coahuila)
[BONAP county map]
Discussion

Euphorbia lurida has been treated as a complex of several taxa in the past, but only a single, broadly-defined species is recognized here. This species is variable in both the pubescence and shape of the bracts subtending the cyathia and also in the degree of crenation of the gland margin. In the northern part of its range, E. lurida appears to intergrade with E. brachycera, and it can be difficult to distinguish these two species in northern Arizona and New Mexico. A report of the species from Sonora, Mexico, based on a single immature collection (V. W. Steinmann and R. S. Felger 1997) has not been verified.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia jejuna is known in the flora area from only a few collections in Mitchell, Nolan, Terrell, and Val Verde counties. The species is very similar to E. astyla, but differs in its more deeply divided involucral gland appendages and more definitely petiolate, rounder leaves.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 305. FNA vol. 12, p. 274.
Parent taxa Euphorbiaceae > Euphorbia > subg. Esula Euphorbiaceae > Euphorbia > sect. Anisophyllum
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms E. palmeri, E. palmeri var. subpubens, E. subpubens, Tithymalus luridus, T. palmeri, T. subpubens Chamaesyce jejuna
Name authority Engelmann: in J. C. Ives, Rep. Colorado R. 4: 26. (1861) M. C. Johnston & Warnock: SouthW. Naturalist 5: 97, fig. [p. 98]. (1960)
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