Euphorbia lurida |
Euphorbia cyparissias |
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woodland spurge |
cypress spurge, euphorbe cypress, graveyard spurge |
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Habit | Herbs, perennial, with thick rootstock. | Herbs, perennial, with slender, spreading rootstock. |
Stems | erect or ascending, unbranched, sometimes sinuous, 5–30 cm, glabrous or sparsely to densely puberulent. |
erect or ascending, unbranched or branched, often with short axillary vegetative shoots with very narrow leaves, 10–50 cm, glabrous. |
Leaves | petiole 0–1 mm; blade oblanceolate to obovate, 8–20 mm × 3–7 mm, base truncate or cuneate, margins entire, apex obtuse to rounded, minutely mucronate, surfaces puberulent or glabrous; venation inconspicuous, only midvein prominent. |
petiole absent; blade linear to linear-ovate or linear-oblanceolate, 5–30 × 0.5–3 mm, base rounded to cuneate, margins entire, apex rounded to acute, surfaces glabrous; venation inconspicuous, only midvein prominent. |
Involucre | cupulate, 2–2.2 × 1.3–1.8 mm, glabrous; glands 4, oblong to broadly ovate, usually truncate, 0.5–0.8 × 1–1.6 mm, margins irregularly crenate to strongly dentate; horns absent or usually divergent or straight, 0.1–0.3 mm, usually slightly longer than, occasionally equaling, teeth on gland margin. |
campanulate to slightly urceolate, 1.5–2 × 0.9–1.1 mm, glabrous; glands 4, crescent-shaped; 0.4–0.6 × 0.7–1.3 mm; horns convergent, 0.1–0.5 mm. |
Staminate flowers | 10–20. |
15–25. |
Pistillate flowers | ovary glabrous or puberulent; styles 0.7–1 mm, 2-fid. |
ovary glabrous; styles 1–1.2 mm, 2-fid. |
Capsules | ovoid, 3.5–4 × 4–4.5 mm, 3-lobed; cocci rounded, smooth to slightly rugose, glabrous; columella 3.2–3.5 mm. |
subglobose, 2.5–3 × 3–4 mm, slightly lobed; cocci rounded, granulate, glabrous; columella 1.9–2.1 mm. |
Seeds | gray to dark gray, truncate-oblong to truncate-ovoid, 2.8–3 × 1.7–2 mm, irregularly pitted; caruncle conic, 0.6 × 0.7 mm. |
blackish, ovoid-oblong, 1.8–2.5 × 1.4–1.7 mm, smooth; caruncle nipple-shaped or subreniform, 0.2–0.4 × 0.7–1.1 mm. |
Cyathial | arrangement: terminal pleiochasial branches 3–5, each 1–4 times 2-branched; pleiochasial bracts ovate to broadly ovate or oblanceolate, wider than distal leaves; dichasial bracts distinct, rounded, oblanceolate, or subreniform, base cuneate or obtuse, margins entire, apex obtuse, slightly mucronate; axillary cymose branches 0–4. |
arrangement: terminal pleiochasial branches 6–25, each 1–2-times 2-branched; pleiochasial bracts similar in shape to distal leaves except shorter and wider; dichasial bracts distinct, widely ovate, rhombic or reniform, base obtuse to truncate, margins entire, apex acute or obtuse, sometimes mucronulate; axillary cymose branches 0–15. |
Cyathia | peduncle 0.3–0.9 mm. |
peduncle 0–0.5 mm. |
Euphorbia lurida |
Euphorbia cyparissias |
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Phenology | Flowering and fruiting spring–summer. | Flowering and fruiting spring–fall. |
Habitat | Open pine-oak forests, dry slopes and canyons. | Fields, roadsides, waste places. |
Elevation | 1300–2800 m. (4300–9200 ft.) | 0–1500 m. (0–4900 ft.) |
Distribution |
AZ; CA; NM; NV; UT; Mexico (Baja California)
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AR; CA; CO; CT; DC; DE; GA; IA; ID; IL; IN; KY; MA; MD; ME; MI; MN; MO; MT; NC; NE; NH; NJ; NY; OH; OR; PA; RI; SC; TN; UT; VA; VT; WA; WI; WV; WY; BC; MB; NB; NF; NS; ON; PE; QC; SK; Europe [Introduced in North America]
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Discussion | Euphorbia lurida has been treated as a complex of several taxa in the past, but only a single, broadly-defined species is recognized here. This species is variable in both the pubescence and shape of the bracts subtending the cyathia and also in the degree of crenation of the gland margin. In the northern part of its range, E. lurida appears to intergrade with E. brachycera, and it can be difficult to distinguish these two species in northern Arizona and New Mexico. A report of the species from Sonora, Mexico, based on a single immature collection (V. W. Steinmann and R. S. Felger 1997) has not been verified. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 12, p. 305. | FNA vol. 12, p. 301. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > subg. Esula |
Sibling taxa | ||
Synonyms | E. palmeri, E. palmeri var. subpubens, E. subpubens, Tithymalus luridus, T. palmeri, T. subpubens | Galarhoeus cyparissias, Tithymalus cyparissias |
Name authority | Engelmann: in J. C. Ives, Rep. Colorado R. 4: 26. (1861) | Linnaeus: Sp. Pl. 1: 461. (1753) |
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