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woodland spurge
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fork spurge
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| Herbs, perennial, with thick rootstock. |
Herbs, annual, with slender, fibrous taproot. |
erect or ascending, unbranched, sometimes sinuous, 5–30 cm, glabrous or sparsely to densely puberulent. |
erect, 20–70 cm, glabrous or with few scattered spreading hairs; branches arcuate, branching appearing dichotomous. |
petiole 0–1 mm; blade oblanceolate to obovate, 8–20 mm × 3–7 mm, base truncate or cuneate, margins entire, apex obtuse to rounded, minutely mucronate, surfaces puberulent or glabrous; venation inconspicuous, only midvein prominent. |
usually alternate, occasionally opposite at proximalmost node; petiole 15–49 mm, glabrous; blade usually ovate, rarely oblong or elliptic, 13–54 × 7–38 mm, base usually rounded to broadly cuneate, rarely truncate, margins finely serrulate, apex obtuse, surfaces glabrous or with few scattered hairs; venation pinnate, midvein prominent. |
cupulate, 2–2.2 × 1.3–1.8 mm, glabrous; glands 4, oblong to broadly ovate, usually truncate, 0.5–0.8 × 1–1.6 mm, margins irregularly crenate to strongly dentate; horns absent or usually divergent or straight, 0.1–0.3 mm, usually slightly longer than, occasionally equaling, teeth on gland margin. |
tubular or obconic, 1–1.7 × 0.7–1.4 mm, glabrous except for few hairs on lobes; involucral lobes divided into several linear, smooth lobes; glands 1(–3), greenish, sessile and broadly attached, 0.3–0.4 × 0.4–0.8 mm, opening oblong to subcircular, glabrous; appendages petaloid, white, elliptic, oblong, transversely oblong, or forming thin, lunate rim on gland margin, not incurved and covering glands, 0.3–0.9 × 0.6–1.3 mm, entire, undulate or slightly lobed, glabrous. |
10–20. |
20–30. |
ovary glabrous or puberulent; styles 0.7–1 mm, 2-fid. |
ovary glabrous; styles 0.6–1 mm, 2-fid 1/2 length. |
ovoid, 3.5–4 × 4–4.5 mm, 3-lobed; cocci rounded, smooth to slightly rugose, glabrous; columella 3.2–3.5 mm. |
oblate, 2.8–3.1 × 3.6–4.5 mm, glabrous; columella 1.9–2.4 mm. |
gray to dark gray, truncate-oblong to truncate-ovoid, 2.8–3 × 1.7–2 mm, irregularly pitted; caruncle conic, 0.6 × 0.7 mm. |
brown to blackish, ovoid, rounded in cross section, 1.9–2.4 × 1.5–1.8 mm, irregularly and coarsely tuberculate; caruncle absent or rudimentary. |
arrangement: terminal pleiochasial branches 3–5, each 1–4 times 2-branched; pleiochasial bracts ovate to broadly ovate or oblanceolate, wider than distal leaves; dichasial bracts distinct, rounded, oblanceolate, or subreniform, base cuneate or obtuse, margins entire, apex obtuse, slightly mucronate; axillary cymose branches 0–4. |
arrangement: terminal dichasial branches 2, few-branched (weakly defined); pleiochasial bracts 2–3, opposite or whorled, wholly green, similar in shape and size to distal leaves; dichasial bracts smaller than distal leaves, often white at base. |
peduncle 0.3–0.9 mm. |
peduncle 0.9–3.5(–6.2) mm. |
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| Flowering and fruiting spring–summer. |
Flowering and fruiting summer–fall. |
| Open pine-oak forests, dry slopes and canyons. |
Riparian areas with cottonwoods and willows, pinyon pine woodlands, pine-oak forests, Douglas fir forests with pines. |
| 1300–2800 m. (4300–9200 ft.) |
1900–2300 m. (6200–7500 ft.) |
|
AZ; CA; NM; NV; UT; Mexico (Baja California)
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NM; TX; Mexico; Central America (Guatemala) |
Euphorbia lurida has been treated as a complex of several taxa in the past, but only a single, broadly-defined species is recognized here. This species is variable in both the pubescence and shape of the bracts subtending the cyathia and also in the degree of crenation of the gland margin. In the northern part of its range, E. lurida appears to intergrade with E. brachycera, and it can be difficult to distinguish these two species in northern Arizona and New Mexico. A report of the species from Sonora, Mexico, based on a single immature collection (V. W. Steinmann and R. S. Felger 1997) has not been verified. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia bifurcata is found in the mountains of southern New Mexico (Doña Ana, Grant, Lincoln, Otero, and Sierra counties) and trans-Pecos Texas (Brewster, Jeff Davis, and Presidio counties). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| FNA vol. 12, p. 305. |
FNA vol. 12, p. 319. |
| Euphorbiaceae > Euphorbia > subg. Esula |
Euphorbiaceae > Euphorbia > sect. Poinsettia |
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana |
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana |
| E. palmeri, E. palmeri var. subpubens, E. subpubens, Tithymalus luridus, T. palmeri, T. subpubens |
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| Engelmann: in J. C. Ives, Rep. Colorado R. 4: 26. (1861) |
Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 190. (1859) |
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