Euphorbia lurida |
Euphorbia austrotexana |
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woodland spurge |
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Habit | Herbs, perennial, with thick rootstock. | Herbs, annual, with taproot. | ||||
Stems | erect or ascending, unbranched, sometimes sinuous, 5–30 cm, glabrous or sparsely to densely puberulent. |
erect, usually branched near base, 6–22 cm, glabrous. |
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Leaves | petiole 0–1 mm; blade oblanceolate to obovate, 8–20 mm × 3–7 mm, base truncate or cuneate, margins entire, apex obtuse to rounded, minutely mucronate, surfaces puberulent or glabrous; venation inconspicuous, only midvein prominent. |
petiole absent; blade linear to slightly lanceolate or linear-oblanceolate, 5–18 × 0.5–2.5 mm, base linear attenuate, margins entire, apex rounded to obtuse or acute, surfaces glabrous; venation pinnate, midvein prominent. |
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Involucre | cupulate, 2–2.2 × 1.3–1.8 mm, glabrous; glands 4, oblong to broadly ovate, usually truncate, 0.5–0.8 × 1–1.6 mm, margins irregularly crenate to strongly dentate; horns absent or usually divergent or straight, 0.1–0.3 mm, usually slightly longer than, occasionally equaling, teeth on gland margin. |
infundibular, 0.8–1.1 × 0.6–0.9 mm, glabrous; glands 4, crescent-shaped; 0.2–0.4 × 0.5–0.6 mm; horns divergent, 0.5–0.7 mm. |
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Staminate flowers | 10–20. |
5–10. |
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Pistillate flowers | ovary glabrous or puberulent; styles 0.7–1 mm, 2-fid. |
ovary glabrous; styles 0.3–0.5 mm, 2-fid. |
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Capsules | ovoid, 3.5–4 × 4–4.5 mm, 3-lobed; cocci rounded, smooth to slightly rugose, glabrous; columella 3.2–3.5 mm. |
ovoid-globose, 1.8–2.2 × 3–3.2 mm, slightly 3-lobed; cocci rounded, smooth, glabrous; columella 1.5–2 mm. |
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Seeds | gray to dark gray, truncate-oblong to truncate-ovoid, 2.8–3 × 1.7–2 mm, irregularly pitted; caruncle conic, 0.6 × 0.7 mm. |
white to gray, ellipsoid, 1.4–1.7 × 1–1.3 mm, with deep, irregular to rounded, shallow to concave depressions over entire surface; caruncle reniform-ovate, depressed-conic, 0.5–0.7 × 0.7–1 mm. |
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Cyathial | arrangement: terminal pleiochasial branches 3–5, each 1–4 times 2-branched; pleiochasial bracts ovate to broadly ovate or oblanceolate, wider than distal leaves; dichasial bracts distinct, rounded, oblanceolate, or subreniform, base cuneate or obtuse, margins entire, apex obtuse, slightly mucronate; axillary cymose branches 0–4. |
arrangement: terminal pleiochasial branches 3, 1–3 times 2-branched; pleiochasial bracts similar in shape to but slightly shorter and wider than distal leaves; dichasial bracts distinct, not imbricate, reniform-ovate to subdeltate-ovate or broadly ovate-lanceolate, base obliquely truncate to rounded, margins entire, apex obtuse to broadly acuminate; axillary cymose branches 0–3. |
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Cyathia | peduncle 0.3–0.9 mm. |
peduncle 0–0.5 mm. |
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Euphorbia lurida |
Euphorbia austrotexana |
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Phenology | Flowering and fruiting spring–summer. | |||||
Habitat | Open pine-oak forests, dry slopes and canyons. | |||||
Elevation | 1300–2800 m. (4300–9200 ft.) | |||||
Distribution |
AZ; CA; NM; NV; UT; Mexico (Baja California)
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TX |
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Discussion | Euphorbia lurida has been treated as a complex of several taxa in the past, but only a single, broadly-defined species is recognized here. This species is variable in both the pubescence and shape of the bracts subtending the cyathia and also in the degree of crenation of the gland margin. In the northern part of its range, E. lurida appears to intergrade with E. brachycera, and it can be difficult to distinguish these two species in northern Arizona and New Mexico. A report of the species from Sonora, Mexico, based on a single immature collection (V. W. Steinmann and R. S. Felger 1997) has not been verified. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). Euphorbia austrotexana occurs in stabilized sandy soil in the south Texas plains (M. H. Mayfield 2013). It is similar to E. longicruris but differs from that species in its often narrowly oblanceolate to linear leaves and its white to gray, ellipsoid seeds that are covered with minute, concave depressions. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 12, p. 305. | FNA vol. 12, p. 298. | ||||
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > subg. Esula | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | E. palmeri, E. palmeri var. subpubens, E. subpubens, Tithymalus luridus, T. palmeri, T. subpubens | |||||
Name authority | Engelmann: in J. C. Ives, Rep. Colorado R. 4: 26. (1861) | Mayfield: J. Bot. Res. Inst. Texas 7: 634, figs. 1, 2[row 3, left & center]. (2013) | ||||
Web links |