Euphorbia lurida |
Euphorbia arizonica |
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woodland spurge |
Arizona sandmat, Arizona spurge |
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Habit | Herbs, perennial, with thick rootstock. | Herbs, annual or short-lived perennial, with slender to slightly thickened taproot. |
Stems | erect or ascending, unbranched, sometimes sinuous, 5–30 cm, glabrous or sparsely to densely puberulent. |
erect to ascending, 10–30 cm, uniformly pilose with glistening hairs. |
Leaves | petiole 0–1 mm; blade oblanceolate to obovate, 8–20 mm × 3–7 mm, base truncate or cuneate, margins entire, apex obtuse to rounded, minutely mucronate, surfaces puberulent or glabrous; venation inconspicuous, only midvein prominent. |
opposite; stipules distinct, deltate, (0–)0.1 mm, glabrous or with few scattered hairs; petiole 0.4–1.5 mm, pilose with glistening hairs; blade usually ovate, rarely elliptic, 3–11 × 2–7 mm, base asymmetric, one side cuneate to rounded, other side rounded to strongly cordate, margins entire, apex obtuse or acute, surfaces pilose with glistening hairs; 3-veined from base but usually only midvein conspicuous. |
Involucre | cupulate, 2–2.2 × 1.3–1.8 mm, glabrous; glands 4, oblong to broadly ovate, usually truncate, 0.5–0.8 × 1–1.6 mm, margins irregularly crenate to strongly dentate; horns absent or usually divergent or straight, 0.1–0.3 mm, usually slightly longer than, occasionally equaling, teeth on gland margin. |
urceolate, 1.1–1.3 × 0.5–0.9 mm, glabrous or pilose with glistening hairs; glands 4, dark maroon, usually oblong to reniform, rarely almost circular, 0.2 × 0.2–0.4 mm; appendages white to pink, oblong, flabellate, oblong, or elliptic, 0.3–0.6 × 0.6–0.9 mm, distal margin entire or crenulate. |
Staminate flowers | 10–20. |
5–12. |
Pistillate flowers | ovary glabrous or puberulent; styles 0.7–1 mm, 2-fid. |
ovary pilose, styles 0.5–0.6 mm, 2-fid 1/2 length. |
Capsules | ovoid, 3.5–4 × 4–4.5 mm, 3-lobed; cocci rounded, smooth to slightly rugose, glabrous; columella 3.2–3.5 mm. |
broadly ovoid to subglobose, 1.4–1.8 mm diam., pilose; columella 1.1–1.4 mm. |
Seeds | gray to dark gray, truncate-oblong to truncate-ovoid, 2.8–3 × 1.7–2 mm, irregularly pitted; caruncle conic, 0.6 × 0.7 mm. |
gray to light brown, ovoid, 4-angled in cross section, 0.9–1.1 × 0.5–0.6 mm, rugose with 2–5 irregular transverse ridges that sometimes pass through abaxial keel. |
Cyathial | arrangement: terminal pleiochasial branches 3–5, each 1–4 times 2-branched; pleiochasial bracts ovate to broadly ovate or oblanceolate, wider than distal leaves; dichasial bracts distinct, rounded, oblanceolate, or subreniform, base cuneate or obtuse, margins entire, apex obtuse, slightly mucronate; axillary cymose branches 0–4. |
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Cyathia | peduncle 0.3–0.9 mm. |
solitary at distal nodes; peduncle 0.7–3.6 mm. |
Euphorbia lurida |
Euphorbia arizonica |
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Phenology | Flowering and fruiting spring–summer. | Flowering and fruiting year-round in response to sufficient moisture. |
Habitat | Open pine-oak forests, dry slopes and canyons. | Washes and rocky slopes, sometimes on limestone, desert scrub communities often with creosote-bush, riparian forests, mesquite woodlands, oak chaparral. |
Elevation | 1300–2800 m. (4300–9200 ft.) | 100–1400 m. (300–4600 ft.) |
Distribution |
AZ; CA; NM; NV; UT; Mexico (Baja California)
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AZ; CA; NM; TX; Mexico (Baja California, Baja California Sur, Chihuahua, Coahuila, Durango, Sonora)
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Discussion | Euphorbia lurida has been treated as a complex of several taxa in the past, but only a single, broadly-defined species is recognized here. This species is variable in both the pubescence and shape of the bracts subtending the cyathia and also in the degree of crenation of the gland margin. In the northern part of its range, E. lurida appears to intergrade with E. brachycera, and it can be difficult to distinguish these two species in northern Arizona and New Mexico. A report of the species from Sonora, Mexico, based on a single immature collection (V. W. Steinmann and R. S. Felger 1997) has not been verified. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia arizonica is distinctive and easily recognized by its glistening, translucent hairs that appear somewhat glutinous and are most apparent on the stems. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 305. | FNA vol. 12, p. 260. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | E. palmeri, E. palmeri var. subpubens, E. subpubens, Tithymalus luridus, T. palmeri, T. subpubens | Chamaesyce arizonica |
Name authority | Engelmann: in J. C. Ives, Rep. Colorado R. 4: 26. (1861) | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 186. (1859) |
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