Euphorbia lurida |
Euphorbia antisyphilitica |
|
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woodland spurge |
candelilla, wax plant |
|
Habit | Herbs, perennial, with thick rootstock. | Shrubs, with much-branched, fleshy rootstock. |
Stems | erect or ascending, unbranched, sometimes sinuous, 5–30 cm, glabrous or sparsely to densely puberulent. |
erect, few branched, 25–50(–100) cm, glabrous or puberulent, pencil-like, in age covered with flaky, exfoliating layer of wax. |
Leaves | petiole 0–1 mm; blade oblanceolate to obovate, 8–20 mm × 3–7 mm, base truncate or cuneate, margins entire, apex obtuse to rounded, minutely mucronate, surfaces puberulent or glabrous; venation inconspicuous, only midvein prominent. |
alternate usually caducous, sometimes persisting; stipules 0.4–0.5 mm; petiole absent; blade ovate to deltate-subulate, 2.5–4 × 1 mm, thick, fleshy, base usually rounded and swollen, rarely cuneate, margins entire, apex acute, surfaces puberulent, adaxial sometimes canescent; venation inconspicuous. |
Involucre | cupulate, 2–2.2 × 1.3–1.8 mm, glabrous; glands 4, oblong to broadly ovate, usually truncate, 0.5–0.8 × 1–1.6 mm, margins irregularly crenate to strongly dentate; horns absent or usually divergent or straight, 0.1–0.3 mm, usually slightly longer than, occasionally equaling, teeth on gland margin. |
campanulate, 1.6–2.2 × 1.6–1.9 mm, puberulent to canescent; glands 5, pinkish, narrowly oblong to reniform, 0.3–0.4 × 0.8–1 mm; appendages white to pink, ovate, oblong, or transversely oblong, 1.3–2.5 × 1.4–2.5 mm, usually erose, rarely entire. |
Staminate flowers | 10–20. |
50–70. |
Pistillate flowers | ovary glabrous or puberulent; styles 0.7–1 mm, 2-fid. |
ovary glabrous; styles 0.9–1.1 mm, 2-fid nearly entire length. |
Capsules | ovoid, 3.5–4 × 4–4.5 mm, 3-lobed; cocci rounded, smooth to slightly rugose, glabrous; columella 3.2–3.5 mm. |
oblong to ovoid, 3.9–4.2 × 3.6–3.9 mm, glabrous; columella 3.1–3.3 mm. |
Seeds | gray to dark gray, truncate-oblong to truncate-ovoid, 2.8–3 × 1.7–2 mm, irregularly pitted; caruncle conic, 0.6 × 0.7 mm. |
whitish gray, narrowly ovoid, 2.4–3.1 × 1.4–1.6 mm, irregularly rugose-tuberculate; caruncle crescent-shaped, 0.3–0.6 × 0.6–0.8 mm. |
Cyathial | arrangement: terminal pleiochasial branches 3–5, each 1–4 times 2-branched; pleiochasial bracts ovate to broadly ovate or oblanceolate, wider than distal leaves; dichasial bracts distinct, rounded, oblanceolate, or subreniform, base cuneate or obtuse, margins entire, apex obtuse, slightly mucronate; axillary cymose branches 0–4. |
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Cyathia | peduncle 0.3–0.9 mm. |
in axillary congested cymes, near branch tips or solitary at distal nodes; peduncle 0–1 mm, lanulose. |
Euphorbia lurida |
Euphorbia antisyphilitica |
|
Phenology | Flowering and fruiting spring–summer. | Flowering and fruiting year-round in response to sufficient rainfall. |
Habitat | Open pine-oak forests, dry slopes and canyons. | Desert scrub, frequently on limestone substrates. |
Elevation | 1300–2800 m. (4300–9200 ft.) | 100–1200 m. (300–3900 ft.) |
Distribution |
AZ; CA; NM; NV; UT; Mexico (Baja California)
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NM; TX; Mexico
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Discussion | Euphorbia lurida has been treated as a complex of several taxa in the past, but only a single, broadly-defined species is recognized here. This species is variable in both the pubescence and shape of the bracts subtending the cyathia and also in the degree of crenation of the gland margin. In the northern part of its range, E. lurida appears to intergrade with E. brachycera, and it can be difficult to distinguish these two species in northern Arizona and New Mexico. A report of the species from Sonora, Mexico, based on a single immature collection (V. W. Steinmann and R. S. Felger 1997) has not been verified. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia antisyphilitica is the only pencil-stemmed species of Euphorbia occurring in the flora area. The species is characteristic of the Chihuahuan Desert scrub of Mexico from Chihuahua and Coahuila south to Hidalgo and Querétaro, and barely enters into the United States in southern New Mexico (Doña Ana and Lincoln counties) and southwest (Brewster, Hudspeth, Presidio, and Terrell counties) and south (Starr and Webb counties) Texas. The stems are covered in a conspicuous coat of exfoliating wax, and the plants historically have been harvested for this product, although the practice is much less prevalent now. The specific epithet refers to its traditional medicinal use in treating sexually transmitted infections. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 305. | FNA vol. 12, p. 243. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Alectoroctonum |
Sibling taxa | ||
Synonyms | E. palmeri, E. palmeri var. subpubens, E. subpubens, Tithymalus luridus, T. palmeri, T. subpubens | |
Name authority | Engelmann: in J. C. Ives, Rep. Colorado R. 4: 26. (1861) | Zuccarini: Flora 15(2, Beibl.): 58. (1832) |
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