Euphorbia lurida |
Euphorbia aaron-rossii |
|
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woodland spurge |
Marble Canyon spurge, Ross' or Marble Canyon spurge |
|
Habit | Herbs, perennial, with thick rootstock. | Herbs, perennial, with deep stout rootstock. |
Stems | erect or ascending, unbranched, sometimes sinuous, 5–30 cm, glabrous or sparsely to densely puberulent. |
erect, branched, densely clumped, previous year’s dead stems persistent, 25–45(–60) cm, glabrous, striate. |
Leaves | petiole 0–1 mm; blade oblanceolate to obovate, 8–20 mm × 3–7 mm, base truncate or cuneate, margins entire, apex obtuse to rounded, minutely mucronate, surfaces puberulent or glabrous; venation inconspicuous, only midvein prominent. |
alternate, persisting, usually reflexed, occasionally spreading; stipules 0.1–0.3 mm; petiole 0.2–2.2 mm, glabrous; blade narrowly ovate to lanceolate proximally, narrowly lanceolate, linear, or filiform distally, 10–32 × 0.5–6.5 mm, base cuneate, margins entire, apex acute, surfaces pilose when young, sparsely strigose or glabrous with age; venation obscure, only midvein conspicuous. |
Involucre | cupulate, 2–2.2 × 1.3–1.8 mm, glabrous; glands 4, oblong to broadly ovate, usually truncate, 0.5–0.8 × 1–1.6 mm, margins irregularly crenate to strongly dentate; horns absent or usually divergent or straight, 0.1–0.3 mm, usually slightly longer than, occasionally equaling, teeth on gland margin. |
turbinate to campanulate, 2.2–3.7 × 1.5–2.5 mm, moderately strigose; glands 5, dark green, reniform, 0.7–1.1 × 1–1.6 mm; appendages white to pink, flabellate, 0.5–1.5 × 0.8–2.2 mm, dentate or erose. |
Staminate flowers | 10–20. |
20–25. |
Pistillate flowers | ovary glabrous or puberulent; styles 0.7–1 mm, 2-fid. |
ovary strigose; styles 1–1.3 mm, 2-fid at apex. |
Capsules | ovoid, 3.5–4 × 4–4.5 mm, 3-lobed; cocci rounded, smooth to slightly rugose, glabrous; columella 3.2–3.5 mm. |
subglobose, 2–3 × 4 mm, sparsely strigose; columella 2–3 mm. |
Seeds | gray to dark gray, truncate-oblong to truncate-ovoid, 2.8–3 × 1.7–2 mm, irregularly pitted; caruncle conic, 0.6 × 0.7 mm. |
gray-green to gray-brown, globose-ovoid, 1.8–2.2 × 1.2–1.6 mm, longitudinally pitted; caruncle absent. |
Cyathial | arrangement: terminal pleiochasial branches 3–5, each 1–4 times 2-branched; pleiochasial bracts ovate to broadly ovate or oblanceolate, wider than distal leaves; dichasial bracts distinct, rounded, oblanceolate, or subreniform, base cuneate or obtuse, margins entire, apex obtuse, slightly mucronate; axillary cymose branches 0–4. |
|
Cyathia | peduncle 0.3–0.9 mm. |
in terminal monochasia (thus appearing solitary at alternate nodes); peduncle 0.5–2.5 (or 10–25) mm, glabrous or sparsely strigose. |
Euphorbia lurida |
Euphorbia aaron-rossii |
|
Phenology | Flowering and fruiting spring–summer. | Flowering and fruiting spring–fall. |
Habitat | Open pine-oak forests, dry slopes and canyons. | Sandy soils and dunes, occasionally rocky slopes, riparian areas. |
Elevation | 1300–2800 m. (4300–9200 ft.) | 600–1300 m. (2000–4300 ft.) |
Distribution |
AZ; CA; NM; NV; UT; Mexico (Baja California)
|
AZ |
Discussion | Euphorbia lurida has been treated as a complex of several taxa in the past, but only a single, broadly-defined species is recognized here. This species is variable in both the pubescence and shape of the bracts subtending the cyathia and also in the degree of crenation of the gland margin. In the northern part of its range, E. lurida appears to intergrade with E. brachycera, and it can be difficult to distinguish these two species in northern Arizona and New Mexico. A report of the species from Sonora, Mexico, based on a single immature collection (V. W. Steinmann and R. S. Felger 1997) has not been verified. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia aaron-rossii is restricted to the banks of the Colorado River in several small areas of the Grand and Marble canyons. The species is most closely related to E. strictior and E. wrightii, but due to its rarity, it has not been extensively studied. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 305. | FNA vol. 12, p. 242. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Alectoroctonum |
Sibling taxa | ||
Synonyms | E. palmeri, E. palmeri var. subpubens, E. subpubens, Tithymalus luridus, T. palmeri, T. subpubens | |
Name authority | Engelmann: in J. C. Ives, Rep. Colorado R. 4: 26. (1861) | A. H. Holmgren & N. H. Holmgren: Brittonia 40: 357, figs. 1, 2. (1988) |
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