Euphorbia lathyris |
Euphorbia theriaca |
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caper spurge, euphorbe épurge, gopher plant, gopher spurge, mole plant |
Terlingua sandmat, Terlingua spurge |
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Habit | Herbs, annual or biennial, with taproot. | Herbs, annual, with slender taproot. | ||||
Stems | erect, unbranched or branched, 200 cm, glabrous, glaucous. |
prostrate to reclining, not mat-forming, 5–30 cm, glabrous. |
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Leaves | opposite, decussate; petiole absent; blade linear to oblong-lanceolate, 30–120 × 3–25 mm, base acute, rounded, cordate or clasping, margins entire, apex acute or subobtuse, sometimes mucronate, surfaces glabrous, abaxial ± glaucous; venation pinnate, midvein prominent. |
opposite; stipules usually distinct, occasionally connate basally on lower side of stem, subulate or scalelike, usually entire, occasionally 2-fid or margin sparsely ciliate, 0.4–1 mm, glabrous; petiole 0.7–1.2(–1.5) mm, glabrous; blade ovate, oblong, orbiculate, or obovate, 2–7.1 × 1–3.5 mm, base slightly asymmetric, rounded, margins entire, often revolute on drying, apex usually rounded, occasionally slightly emarginate, surfaces glabrous; venation usually obscure, only midvein conspicuous. |
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Involucre | campanulate, 1.2–2.3 × 1.4–2.5 mm, glabrous; glands 4, elliptic, 0.3–0.6 × 1–1.3 mm; horns divergent, thick, tips rounded, dilated, 0.5–1.4 mm. |
usually turbinate-campanulate to hemispheric, occasionally suburceolate, 1–1.8 × 0.9–1.4 mm, glabrous; glands 4, yellow-green to red-purple, sessile or short-stipitate, subcircular to slightly elliptic, 0.2–0.5 × 0.2–0.7 mm; appendages absent or white to pink, semilunate or forming rim at edge of gland, (0–)0.1–0.4 × (0–)0.3–0.9 mm, entire or slightly crenate. |
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Staminate flowers | 25–30. |
15–36. |
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Pistillate flowers | ovary glabrous; styles 0.7–2.2 mm, 2-fid. |
ovary glabrous; styles 0.3–0.5 mm, 2-fid 1/2 length. |
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Capsules | depressed-globose, 9–12 × 12–16 mm, deeply 3-lobed, tardily dehiscent and appearing indehiscent, mesocarp spongy; cocci rounded, smooth, glabrous; columella 4–5.2 mm. |
broadly ovoid, 1.1–1.6 × 1.5–1.8 mm, glabrous; columella 1.2–1.5 mm. |
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Seeds | brownish or blackish, oblong, 4.5–6 × 3–4.2 mm, rugose, irregularly reticulate; caruncle substipitate, hat-shaped, 1.6–2 × 1.2–1.5 mm. |
whitish, reddish brown beneath coat, ovate, 4-angled in cross section, 0.8–1.2 × 0.5–0.8 mm, with (2–)3(–5) deep transverse ridges. |
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Cyathial | arrangement: terminal pleiochasial branches 2–4, each 1–2 times 2-branched; pleiochasial bracts cordate-lanceolate, shorter and wider than distal leaves; dichasial bracts distinct, ovate-oblong to lanceolate, base subcordate, margins entire, apex acute; axillary cymose branches 0–10. |
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Cyathia | peduncle 0–0.5 mm. |
usually solitary at distal nodes, rarely clustered on short, axillary branches; peduncle 0.3–1.3 mm. |
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2n | = 28. |
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Euphorbia lathyris |
Euphorbia theriaca |
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Phenology | Flowering and fruiting winter–fall. | |||||
Habitat | Roadsides, cultivated fields, stream banks, waste places. | |||||
Elevation | 0–1800 m. (0–5900 ft.) | |||||
Distribution |
AZ; CA; CT; ID; IL; MA; MD; MT; NC; OH; OR; PA; SC; TN; TX; VA; WA; WV; BC; ON; QC; Europe; Asia; Africa [Introduced in North America; introduced also in Mexico, South America, Australia]
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NM; TX; n Mexico
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Discussion | Euphorbia lathyris is most likely native to the central and eastern Mediterranean region, but it is widely cultivated and often locally escaped in temperate regions worldwide, as in the flora area. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 12, p. 304. | FNA vol. 12, p. 290. | ||||
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Anisophyllum | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Galarhoeus lathyris, Tithymalus lathyris | Chamaesyce theriaca | ||||
Name authority | Linnaeus: Sp. Pl. 1: 457. (1753) — (as lathyrus) | L. C. Wheeler: Rhodora 43: 242, plate 660, fig. A. (1941) | ||||
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