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caper spurge, euphorbe épurge, gopher plant, gopher spurge, mole plant

weak spurge

Habit Herbs, annual or biennial, with taproot. Herbs, annual, with taproot.
Stems

erect, unbranched or branched, 200 cm, glabrous, glaucous.

erect, unbranched, 7–20 cm, glabrous.

Leaves

opposite, decussate;

petiole absent;

blade linear to oblong-lanceolate, 30–120 × 3–25 mm, base acute, rounded, cordate or clasping, margins entire, apex acute or subobtuse, sometimes mucronate, surfaces glabrous, abaxial ± glaucous;

venation pinnate, midvein prominent.

petiole 1–2 mm, reduced distally;

blade spatulate-cuneate, 8–10 × 4–5 mm (greatly reduced in size proximally), base cuneate, margins entire, apex rounded to emarginate or obcordate, surfaces glabrous;

venation pinnate.

Involucre

campanulate, 1.2–2.3 × 1.4–2.5 mm, glabrous;

glands 4, elliptic, 0.3–0.6 × 1–1.3 mm;

horns divergent, thick, tips rounded, dilated, 0.5–1.4 mm.

campanulate, 0.8–1.1 × 0.7–1 mm, glabrous;

glands 4, elliptic to trapezoidal, 0.3–0.6 × 0.6–1.2 mm;

horns divergent, 0.5–1 mm.

Staminate flowers

25–30.

10–15.

Pistillate flowers

ovary glabrous;

styles 0.7–2.2 mm, 2-fid.

ovary smooth, glabrous;

styles 0.6–1 mm, 2-fid.

Capsules

depressed-globose, 9–12 × 12–16 mm, deeply 3-lobed, tardily dehiscent and appearing indehiscent, mesocarp spongy;

cocci rounded, smooth, glabrous;

columella 4–5.2 mm.

depressed-globose, 1.8–2.2 × 2.2–2.9 mm, slightly lobed;

cocci rounded to slightly flattened, smooth, glabrous;

columella 1.5–1.8 mm.

Seeds

brownish or blackish, oblong, 4.5–6 × 3–4.2 mm, rugose, irregularly reticulate;

caruncle substipitate, hat-shaped, 1.6–2 × 1.2–1.5 mm.

reddish brown to brown, often glaucous, oblong, 1.3–1.4 × 0.8–0.9 mm, abaxial faces with 15–20 shallow pits or almost smooth, adaxial faces with 4–6 large shallow pits or irregular oblong grooves;

caruncle conic, hat-shaped, 0.3–0.4 × 0.4–0.6 mm.

Cyathial

arrangement: terminal pleiochasial branches 2–4, each 1–2 times 2-branched;

pleiochasial bracts cordate-lanceolate, shorter and wider than distal leaves;

dichasial bracts distinct, ovate-oblong to lanceolate, base subcordate, margins entire, apex acute;

axillary cymose branches 0–10.

arrangement: terminal pleiochasial branches 3, each 1–3(–4) times 2-branched;

pleiochasial bracts obovate, similar in size to distal leaves;

dichasial bracts distinct or basally subconnate, not imbricate, triangular-ovate, base truncate or cordate, margins entire, apex mucronate;

axillary cymose branches 1–4.

Cyathia

peduncle 0–0.5 mm.

peduncle 0.2–0.6 mm.

Euphorbia lathyris

Euphorbia tetrapora

Phenology Flowering and fruiting winter–fall. Flowering and fruiting spring.
Habitat Roadsides, cultivated fields, stream banks, waste places. Sandy soils, dry open woods.
Elevation 0–1800 m. (0–5900 ft.) 0–300 m. (0–1000 ft.)
Distribution
from FNA
AZ; CA; CT; ID; IL; MA; MD; MT; NC; OH; OR; PA; SC; TN; TX; VA; WA; WV; BC; ON; QC; Europe; Asia; Africa [Introduced in North America; introduced also in Mexico, South America, Australia]
[WildflowerSearch map]
[BONAP county map]
from FNA
LA; OK; TX
[BONAP county map]
Discussion

Euphorbia lathyris is most likely native to the central and eastern Mediterranean region, but it is widely cultivated and often locally escaped in temperate regions worldwide, as in the flora area.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia tetrapora is endemic to a portion of the western Gulf coastal plain. D. S. Correll and M. C. Johnston (1970) included Alabama and Georgia in the distribution of this species as well, probably due to Engelmann’s citation of a Georgia specimen from the herbarium of Samuel Boykin. Whether the Boykin specimen came from Georgia, where Boykin was based, is unclear. Because no records to support its occurrence in the eastern Gulf coastal plain (Alabama or Georgia) have been found, those states are here excluded from the distribution of E. tetrapora.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 304. FNA vol. 12, p. 311.
Parent taxa Euphorbiaceae > Euphorbia > subg. Esula Euphorbiaceae > Euphorbia > subg. Esula
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms Galarhoeus lathyris, Tithymalus lathyris Tithymalus tetraporus
Name authority Linnaeus: Sp. Pl. 1: 457. (1753) — (as lathyrus) Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 191. (1859)
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