Euphorbia lathyris |
Euphorbia stictospora |
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caper spurge, euphorbe épurge, gopher plant, gopher spurge, mole plant |
mat spurge, narrow-seed spurge, slimseed sandmat |
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Habit | Herbs, annual or biennial, with taproot. | Herbs, annual, with taproot. |
Stems | erect, unbranched or branched, 200 cm, glabrous, glaucous. |
prostrate, often mat-forming, occasionally with ascending tips, 5–45 cm, densely and evenly pilose to lanate. |
Leaves | opposite, decussate; petiole absent; blade linear to oblong-lanceolate, 30–120 × 3–25 mm, base acute, rounded, cordate or clasping, margins entire, apex acute or subobtuse, sometimes mucronate, surfaces glabrous, abaxial ± glaucous; venation pinnate, midvein prominent. |
opposite; stipules distinct or connate basally on one side of stem, entire or irregularly toothed or fringed, 0.5–1.2 mm, pilose to lanate; petiole 0.3–1.5 mm, pilose to lanate; blade usually oblong to oblong-obovate, occasionally nearly circular, 3–10(–15) × 2–5(–10) mm, base asymmetric, one side usually angled or rounded and other truncate-auriculate, margins minutely or conspicuously serrulate at least toward apex, apex usually broadly rounded to broadly acute, occasionally emarginate, abaxial surface often ± lighter green and without reddish spot, both surfaces sparsely to moderately pilose to lanate; 3-veined from base or venation obscure. |
Involucre | campanulate, 1.2–2.3 × 1.4–2.5 mm, glabrous; glands 4, elliptic, 0.3–0.6 × 1–1.3 mm; horns divergent, thick, tips rounded, dilated, 0.5–1.4 mm. |
obconic, 0.7–1 × 0.4–0.6 mm, moderately to densely pilose to lanate; glands 4, reddish, ± unequal, oblong, 0.1 × 0.1–0.3 mm; appendages white to strongly pinkish or reddish tinged, often unequal, sometimes 1 to all absent, 0–0.3 × 0–0.4 mm, 3-lobed or rudimentarily 1-lobed, distal margin crenate. |
Staminate flowers | 25–30. |
3–9. |
Pistillate flowers | ovary glabrous; styles 0.7–2.2 mm, 2-fid. |
ovary pilose to villous, hairs occasionally slightly appressed; styles 0.2–0.5 mm, unbranched or inconspicuously notched at tip. |
Capsules | depressed-globose, 9–12 × 12–16 mm, deeply 3-lobed, tardily dehiscent and appearing indehiscent, mesocarp spongy; cocci rounded, smooth, glabrous; columella 4–5.2 mm. |
ovoid, 1.6–2.3 × 1.4–1.5 mm, moderately to densely villous with hairs usually slightly appressed, pubescence often concentrated on proximal 1/2 or along lobes; columella 1.5–2 mm. |
Seeds | brownish or blackish, oblong, 4.5–6 × 3–4.2 mm, rugose, irregularly reticulate; caruncle substipitate, hat-shaped, 1.6–2 × 1.2–1.5 mm. |
light to dark brown, usually mottled, sometimes with thin, white coating, often wearing away irregularly, narrowly oblong-ovoid to ellipsoid, 3–4-angled in cross section, 1–1.5 × 0.5–0.6 mm, with short, irregularly interrupted furrows, appearing partially and irregularly few-ridged. |
Cyathial | arrangement: terminal pleiochasial branches 2–4, each 1–2 times 2-branched; pleiochasial bracts cordate-lanceolate, shorter and wider than distal leaves; dichasial bracts distinct, ovate-oblong to lanceolate, base subcordate, margins entire, apex acute; axillary cymose branches 0–10. |
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Cyathia | peduncle 0–0.5 mm. |
solitary at leaf nodes or in small, cymose clusters on congested, axillary branches; peduncle 0.7–2.5 mm. |
Euphorbia lathyris |
Euphorbia stictospora |
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Phenology | Flowering and fruiting winter–fall. | Flowering and fruiting midsummer–early fall. |
Habitat | Roadsides, cultivated fields, stream banks, waste places. | Open disturbed areas, rocky slopes. |
Elevation | 0–1800 m. (0–5900 ft.) | 100–2100 m. (300–6900 ft.) |
Distribution |
AZ; CA; CT; ID; IL; MA; MD; MT; NC; OH; OR; PA; SC; TN; TX; VA; WA; WV; BC; ON; QC; Europe; Asia; Africa [Introduced in North America; introduced also in Mexico, South America, Australia]
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AZ; CO; IA; KS; MO; ND; NE; NM; OK; SD; TX; WY; Mexico (Chihuahua, Durango, San Luis Potosí)
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Discussion | Euphorbia lathyris is most likely native to the central and eastern Mediterranean region, but it is widely cultivated and often locally escaped in temperate regions worldwide, as in the flora area. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia stictospora has been recorded from New York, but this disjunct occurrence likely represents a waif or misidentification. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 304. | FNA vol. 12, p. 289. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Galarhoeus lathyris, Tithymalus lathyris | Chamaesyce stictospora |
Name authority | Linnaeus: Sp. Pl. 1: 457. (1753) — (as lathyrus) | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 187. (1859) |
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