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caper spurge, euphorbe épurge, gopher plant, gopher spurge, mole plant

dune spurge, Parry's sandmat, Parry's spurge

Habit Herbs, annual or biennial, with taproot. Herbs, annual, with taproot.
Stems

erect, unbranched or branched, 200 cm, glabrous, glaucous.

usually prostrate, rarely ascending-erect, 5–70(–85) cm, glabrous.

Leaves

opposite, decussate;

petiole absent;

blade linear to oblong-lanceolate, 30–120 × 3–25 mm, base acute, rounded, cordate or clasping, margins entire, apex acute or subobtuse, sometimes mucronate, surfaces glabrous, abaxial ± glaucous;

venation pinnate, midvein prominent.

opposite;

stipules distinct, linear-subulate, usually lacerate and divided into 2 or more slender segments, rarely entire, 0.6–1.4 mm, glabrous;

petiole 1–2.5 mm, glabrous;

blade linear to narrowly oblong, (5–)10–25(–30) × 2–5 mm, base usually symmetric, sometimes slightly asymmetric, attenuate, margins entire, occasionally ± revolute, apex acute to obtuse, mucronulate, surfaces glabrous;

only midvein conspicuous.

Involucre

campanulate, 1.2–2.3 × 1.4–2.5 mm, glabrous;

glands 4, elliptic, 0.3–0.6 × 1–1.3 mm;

horns divergent, thick, tips rounded, dilated, 0.5–1.4 mm.

broadly cupuliform-campanulate, 1.2–1.7 × 1.4–1.8 mm, glabrous;

glands 4, reddish pink to greenish yellow, deeply concave, elliptic to oblong, 0.2–0.3 × 0.3–0.5 mm;

appendages white, elliptic to oblong, usually forming narrow margin around gland, sometimes rudimentary, 0.2–0.6 × 0.3–0.7(–1.1) mm, distal margin entire.

Staminate flowers

25–30.

40–55.

Pistillate flowers

ovary glabrous;

styles 0.7–2.2 mm, 2-fid.

ovary glabrous;

styles 0.5–0.7 mm, 2-fid nearly entire length.

Capsules

depressed-globose, 9–12 × 12–16 mm, deeply 3-lobed, tardily dehiscent and appearing indehiscent, mesocarp spongy;

cocci rounded, smooth, glabrous;

columella 4–5.2 mm.

ovoid-globose, 2–2.3 × 1.5–2.5 mm, glabrous;

columella 1.4–2 mm.

Seeds

brownish or blackish, oblong, 4.5–6 × 3–4.2 mm, rugose, irregularly reticulate;

caruncle substipitate, hat-shaped, 1.6–2 × 1.2–1.5 mm.

mottled brown and white because of irregularly loose and tight outer covering, broadly ovoid, rounded-angular in cross section, 1.4–1.8 × 0.8–1 mm, smooth or only inconspicuously roughened.

Cyathial

arrangement: terminal pleiochasial branches 2–4, each 1–2 times 2-branched;

pleiochasial bracts cordate-lanceolate, shorter and wider than distal leaves;

dichasial bracts distinct, ovate-oblong to lanceolate, base subcordate, margins entire, apex acute;

axillary cymose branches 0–10.

Cyathia

peduncle 0–0.5 mm.

solitary or in small clusters on short axillary branches at distal nodes;

peduncle 1–5 mm.

Euphorbia lathyris

Euphorbia parryi

Phenology Flowering and fruiting winter–fall. Flowering and fruiting spring–fall.
Habitat Roadsides, cultivated fields, stream banks, waste places. Sand dunes, other sandy habitats.
Elevation 0–1800 m. (0–5900 ft.) 200–2200 m. (700–7200 ft.)
Distribution
from FNA
AZ; CA; CT; ID; IL; MA; MD; MT; NC; OH; OR; PA; SC; TN; TX; VA; WA; WV; BC; ON; QC; Europe; Asia; Africa [Introduced in North America; introduced also in Mexico, South America, Australia]
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; NM; NV; TX; UT; Mexico (Chihuahua)
[WildflowerSearch map]
[BONAP county map]
Discussion

Euphorbia lathyris is most likely native to the central and eastern Mediterranean region, but it is widely cultivated and often locally escaped in temperate regions worldwide, as in the flora area.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia parryi is similar to E. missurica, differing only by the generally narrow involucral gland appendages and prostrate habit in E. parryi as opposed to the conspicuous involucral gland appendages and ascending-erect habit in E. missurica. Euphorbia parryi has sometimes been considered the western race of E. missurica (D. S. Correll and M. C. Johnston 1970).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 304. FNA vol. 12, p. 281.
Parent taxa Euphorbiaceae > Euphorbia > subg. Esula Euphorbiaceae > Euphorbia > sect. Anisophyllum
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms Galarhoeus lathyris, Tithymalus lathyris Chamaesyce longeramosa, C. parryi, E. longeramosa
Name authority Linnaeus: Sp. Pl. 1: 457. (1753) — (as lathyrus) Engelmann: Amer. Naturalist 9: 350. (1875)
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