Euphorbia lathyris |
Euphorbia lurida |
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caper spurge, euphorbe épurge, gopher plant, gopher spurge, mole plant |
woodland spurge |
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Habit | Herbs, annual or biennial, with taproot. | Herbs, perennial, with thick rootstock. |
Stems | erect, unbranched or branched, 200 cm, glabrous, glaucous. |
erect or ascending, unbranched, sometimes sinuous, 5–30 cm, glabrous or sparsely to densely puberulent. |
Leaves | opposite, decussate; petiole absent; blade linear to oblong-lanceolate, 30–120 × 3–25 mm, base acute, rounded, cordate or clasping, margins entire, apex acute or subobtuse, sometimes mucronate, surfaces glabrous, abaxial ± glaucous; venation pinnate, midvein prominent. |
petiole 0–1 mm; blade oblanceolate to obovate, 8–20 mm × 3–7 mm, base truncate or cuneate, margins entire, apex obtuse to rounded, minutely mucronate, surfaces puberulent or glabrous; venation inconspicuous, only midvein prominent. |
Involucre | campanulate, 1.2–2.3 × 1.4–2.5 mm, glabrous; glands 4, elliptic, 0.3–0.6 × 1–1.3 mm; horns divergent, thick, tips rounded, dilated, 0.5–1.4 mm. |
cupulate, 2–2.2 × 1.3–1.8 mm, glabrous; glands 4, oblong to broadly ovate, usually truncate, 0.5–0.8 × 1–1.6 mm, margins irregularly crenate to strongly dentate; horns absent or usually divergent or straight, 0.1–0.3 mm, usually slightly longer than, occasionally equaling, teeth on gland margin. |
Staminate flowers | 25–30. |
10–20. |
Pistillate flowers | ovary glabrous; styles 0.7–2.2 mm, 2-fid. |
ovary glabrous or puberulent; styles 0.7–1 mm, 2-fid. |
Capsules | depressed-globose, 9–12 × 12–16 mm, deeply 3-lobed, tardily dehiscent and appearing indehiscent, mesocarp spongy; cocci rounded, smooth, glabrous; columella 4–5.2 mm. |
ovoid, 3.5–4 × 4–4.5 mm, 3-lobed; cocci rounded, smooth to slightly rugose, glabrous; columella 3.2–3.5 mm. |
Seeds | brownish or blackish, oblong, 4.5–6 × 3–4.2 mm, rugose, irregularly reticulate; caruncle substipitate, hat-shaped, 1.6–2 × 1.2–1.5 mm. |
gray to dark gray, truncate-oblong to truncate-ovoid, 2.8–3 × 1.7–2 mm, irregularly pitted; caruncle conic, 0.6 × 0.7 mm. |
Cyathial | arrangement: terminal pleiochasial branches 2–4, each 1–2 times 2-branched; pleiochasial bracts cordate-lanceolate, shorter and wider than distal leaves; dichasial bracts distinct, ovate-oblong to lanceolate, base subcordate, margins entire, apex acute; axillary cymose branches 0–10. |
arrangement: terminal pleiochasial branches 3–5, each 1–4 times 2-branched; pleiochasial bracts ovate to broadly ovate or oblanceolate, wider than distal leaves; dichasial bracts distinct, rounded, oblanceolate, or subreniform, base cuneate or obtuse, margins entire, apex obtuse, slightly mucronate; axillary cymose branches 0–4. |
Cyathia | peduncle 0–0.5 mm. |
peduncle 0.3–0.9 mm. |
Euphorbia lathyris |
Euphorbia lurida |
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Phenology | Flowering and fruiting winter–fall. | Flowering and fruiting spring–summer. |
Habitat | Roadsides, cultivated fields, stream banks, waste places. | Open pine-oak forests, dry slopes and canyons. |
Elevation | 0–1800 m. (0–5900 ft.) | 1300–2800 m. (4300–9200 ft.) |
Distribution |
AZ; CA; CT; ID; IL; MA; MD; MT; NC; OH; OR; PA; SC; TN; TX; VA; WA; WV; BC; ON; QC; Europe; Asia; Africa [Introduced in North America; introduced also in Mexico, South America, Australia]
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AZ; CA; NM; NV; UT; Mexico (Baja California)
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Discussion | Euphorbia lathyris is most likely native to the central and eastern Mediterranean region, but it is widely cultivated and often locally escaped in temperate regions worldwide, as in the flora area. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia lurida has been treated as a complex of several taxa in the past, but only a single, broadly-defined species is recognized here. This species is variable in both the pubescence and shape of the bracts subtending the cyathia and also in the degree of crenation of the gland margin. In the northern part of its range, E. lurida appears to intergrade with E. brachycera, and it can be difficult to distinguish these two species in northern Arizona and New Mexico. A report of the species from Sonora, Mexico, based on a single immature collection (V. W. Steinmann and R. S. Felger 1997) has not been verified. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 304. | FNA vol. 12, p. 305. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > subg. Esula |
Sibling taxa | ||
Synonyms | Galarhoeus lathyris, Tithymalus lathyris | E. palmeri, E. palmeri var. subpubens, E. subpubens, Tithymalus luridus, T. palmeri, T. subpubens |
Name authority | Linnaeus: Sp. Pl. 1: 457. (1753) — (as lathyrus) | Engelmann: in J. C. Ives, Rep. Colorado R. 4: 26. (1861) |
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