Euphorbia lathyris |
Euphorbia fendleri |
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caper spurge, euphorbe épurge, gopher plant, gopher spurge, mole plant |
Fendler's sandmat |
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Habit | Herbs, annual or biennial, with taproot. | Herbs, perennial, with woody, thickened taproot. |
Stems | erect, unbranched or branched, 200 cm, glabrous, glaucous. |
usually prostrate, decumbent, or ascending, very rarely erect, often densely clustered from top of woody crown, 5–12 cm, glabrous. |
Leaves | opposite, decussate; petiole absent; blade linear to oblong-lanceolate, 30–120 × 3–25 mm, base acute, rounded, cordate or clasping, margins entire, apex acute or subobtuse, sometimes mucronate, surfaces glabrous, abaxial ± glaucous; venation pinnate, midvein prominent. |
usually opposite, rarely whorled; stipules distinct, narrowly linear, 0.5–1 mm, glabrous; petiole 0.5–1 mm, glabrous; blade usually orbiculate to ovate, rarely almost lanceolate, 3–8 × 2.5–7 mm, base slightly asymmetric, slightly cordate to rounded or obtuse, margins entire, apex rounded to obtuse, surfaces glabrous; obscurely 3–5-veined from base, only midvein conspicuous. |
Involucre | campanulate, 1.2–2.3 × 1.4–2.5 mm, glabrous; glands 4, elliptic, 0.3–0.6 × 1–1.3 mm; horns divergent, thick, tips rounded, dilated, 0.5–1.4 mm. |
campanulate to turbinate or broadly cupulate, 1.1–1.7 × 1.2–1.8 mm, glabrous; glands 4, yellow-green to reddish, elliptic to oblong, 0.2–0.5 × 0.4–0.9 mm; appendages absent or white, rarely pink, often unequal, lunate to flabellate or sometimes forming crenate margin along gland, (0–)0.1–0.6 × (0–)0.5–1.5 mm, distal margin entire or toothed. |
Staminate flowers | 25–30. |
25–35. |
Pistillate flowers | ovary glabrous; styles 0.7–2.2 mm, 2-fid. |
ovary glabrous; styles 0.3–0.4 mm, 2-fid 1/2 length. |
Capsules | depressed-globose, 9–12 × 12–16 mm, deeply 3-lobed, tardily dehiscent and appearing indehiscent, mesocarp spongy; cocci rounded, smooth, glabrous; columella 4–5.2 mm. |
depressed-globose, 2–2.4 × 2.2–2.5 mm, glabrous; columella 1.7–2.1 mm. |
Seeds | brownish or blackish, oblong, 4.5–6 × 3–4.2 mm, rugose, irregularly reticulate; caruncle substipitate, hat-shaped, 1.6–2 × 1.2–1.5 mm. |
white, ovoid-pyramidal, prominently 4-angled in cross section, 1.7–2 × 1–1.2 mm, smooth to slightly wrinkled. |
Cyathial | arrangement: terminal pleiochasial branches 2–4, each 1–2 times 2-branched; pleiochasial bracts cordate-lanceolate, shorter and wider than distal leaves; dichasial bracts distinct, ovate-oblong to lanceolate, base subcordate, margins entire, apex acute; axillary cymose branches 0–10. |
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Cyathia | peduncle 0–0.5 mm. |
solitary at distal nodes; peduncle 0.7–1.2 mm. |
2n | = 28. |
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Euphorbia lathyris |
Euphorbia fendleri |
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Phenology | Flowering and fruiting winter–fall. | Flowering and fruiting spring–fall. |
Habitat | Roadsides, cultivated fields, stream banks, waste places. | Mountain slopes, desert scrub, pinyon-juniper woodlands, hills, canyons, grasslands, washes, roadsides, dry crevices in limestone, often in gravel and sand. |
Elevation | 0–1800 m. (0–5900 ft.) | 500–2600 m. (1600–8500 ft.) |
Distribution |
AZ; CA; CT; ID; IL; MA; MD; MT; NC; OH; OR; PA; SC; TN; TX; VA; WA; WV; BC; ON; QC; Europe; Asia; Africa [Introduced in North America; introduced also in Mexico, South America, Australia]
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AZ; CA; CO; KS; NE; NM; NV; OK; SD; TX; UT; WY; Mexico (Coahuila)
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Discussion | Euphorbia lathyris is most likely native to the central and eastern Mediterranean region, but it is widely cultivated and often locally escaped in temperate regions worldwide, as in the flora area. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia fendleri is similar to E. chaetocalyx and may sometimes be confused with that species. Its prostrate to decumbent or ascending stems and small, ovate to orbiculate leaves distinguish it from E. chaetocalyx. Some authors have used the presence or absence and shape of the involucral gland appendages to help separate E. fendleri from E. chaetocalyx, but those characters appear highly variable and of little taxonomic utility; somewhat intermediate individuals occur in western Arizona, New Mexico, and Texas. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 304. | FNA vol. 12, p. 267. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Galarhoeus lathyris, Tithymalus lathyris | Chamaesyce fendleri |
Name authority | Linnaeus: Sp. Pl. 1: 457. (1753) — (as lathyrus) | Torrey & A. Gray: in War Department [U.S.], Pacif. Railr. Rep. 2(4): 175. (1857) |
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