Euphorbia lathyris |
Euphorbia exstipulata |
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caper spurge, euphorbe épurge, gopher plant, gopher spurge, mole plant |
square-seed spurge, squareseed or Clark Mountain spurge |
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Habit | Herbs, annual or biennial, with taproot. | Herbs, annual, with slender taproot. |
Stems | erect, unbranched or branched, 200 cm, glabrous, glaucous. |
erect, 5–26 cm, uniformly puberulent to hispidulous or glabrous; branches arcuate. |
Leaves | opposite, decussate; petiole absent; blade linear to oblong-lanceolate, 30–120 × 3–25 mm, base acute, rounded, cordate or clasping, margins entire, apex acute or subobtuse, sometimes mucronate, surfaces glabrous, abaxial ± glaucous; venation pinnate, midvein prominent. |
opposite; petiole 1–3 mm, often indistinct, glabrous or puberulent; blade linear to narrowly elliptic or ovate, 14–42 × 3–28 mm, base attenuate, margins coarsely serrate, occasionally revolute, apex acute to obtuse, abaxial surface sparsely hispidulous to strigillose, adaxial surface glabrous; midvein conspicuous. |
Involucre | campanulate, 1.2–2.3 × 1.4–2.5 mm, glabrous; glands 4, elliptic, 0.3–0.6 × 1–1.3 mm; horns divergent, thick, tips rounded, dilated, 0.5–1.4 mm. |
turbinate to campanulate, 1.1–1.5 × 1–1.3 mm, glabrous, pilose or puberulent; involucral lobes divided into several linear lobes; glands 4(–5), yellow to pink, sessile and broadly attached, 0.2 × 0.3–0.4 mm, opening oblong to nearly circular, glabrous; appendages usually petaloid, white to pink, ovate to trapezoidal, occasionally absent, not incurved and covering glands, 0.2–0.4 × 0.3–0.8 mm, entire, undulate, or conspicuously divided into triangular segments, glabrous. |
Staminate flowers | 25–30. |
10–12. |
Pistillate flowers | ovary glabrous; styles 0.7–2.2 mm, 2-fid. |
ovary puberulent on keels, styles 0.8–1.1 mm, 2-fid 1/2 to nearly entire length. |
Capsules | depressed-globose, 9–12 × 12–16 mm, deeply 3-lobed, tardily dehiscent and appearing indehiscent, mesocarp spongy; cocci rounded, smooth, glabrous; columella 4–5.2 mm. |
broadly depressed-oblong to ovoid, 2.7–3.3 × 3.1–3.9 mm, puberulent (with appressed hairs usually concentrated on keels); columella 1.9–2.5 mm. |
Seeds | brownish or blackish, oblong, 4.5–6 × 3–4.2 mm, rugose, irregularly reticulate; caruncle substipitate, hat-shaped, 1.6–2 × 1.2–1.5 mm. |
white to gray or light brown, ovoid, bluntly 4-angled in cross section, 1.9–2.5 × 1.4–1.7 mm, tuberculate, often with 2 transverse ridges; caruncle 0.1 × 0.2 mm. |
Cyathial | arrangement: terminal pleiochasial branches 2–4, each 1–2 times 2-branched; pleiochasial bracts cordate-lanceolate, shorter and wider than distal leaves; dichasial bracts distinct, ovate-oblong to lanceolate, base subcordate, margins entire, apex acute; axillary cymose branches 0–10. |
arrangement: terminal cymose or dichasial branches usually 1–2, occasionally reduced to monochasia, 1–2-branched; pleiochasial bracts 2–4, often whorled, wholly green or paler green at base, similar in shape and size to distal leaves or slightly narrower; dichasial bracts similar in shape to distal leaves but smaller or highly reduced. |
Cyathia | peduncle 0–0.5 mm. |
peduncle 1–1.9 mm. |
Euphorbia lathyris |
Euphorbia exstipulata |
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Phenology | Flowering and fruiting winter–fall. | Flowering and fruiting summer–fall. |
Habitat | Roadsides, cultivated fields, stream banks, waste places. | Desert scrub, grasslands, mesquite savannas, oak and oak-juniper woodlands. |
Elevation | 0–1800 m. (0–5900 ft.) | 800–2000 m. (2600–6600 ft.) |
Distribution |
AZ; CA; CT; ID; IL; MA; MD; MT; NC; OH; OR; PA; SC; TN; TX; VA; WA; WV; BC; ON; QC; Europe; Asia; Africa [Introduced in North America; introduced also in Mexico, South America, Australia]
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AZ; CA; NM; TX; UT; WY; Mexico (Chihuahua, Coahuila, Durango, Guanajuato, San Luis Potosí, Sonora, Zacatecas)
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Discussion | Euphorbia lathyris is most likely native to the central and eastern Mediterranean region, but it is widely cultivated and often locally escaped in temperate regions worldwide, as in the flora area. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia exstipulata is native from Texas to California and northern Mexico. The species was found once in the late nineteenth century in Wyoming but has not been re-collected there. Broad-leaved plants have been segregated as var. lata, but the variation in leaf shape is continuous and no varieties are formally recognized here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 304. | FNA vol. 12, p. 322. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Poinsettia |
Sibling taxa | ||
Synonyms | Galarhoeus lathyris, Tithymalus lathyris | E. exstipulata var. lata |
Name authority | Linnaeus: Sp. Pl. 1: 457. (1753) — (as lathyrus) | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 189. (1859) |
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