Euphorbia lathyris |
Euphorbia carunculata |
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caper spurge, euphorbe épurge, gopher plant, gopher spurge, mole plant |
sand-dune sandmat or spurge |
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Habit | Herbs, annual or biennial, with taproot. | Herbs, annual, with taproot. |
Stems | erect, unbranched or branched, 200 cm, glabrous, glaucous. |
prostrate, spreading and lanky or occasionally mat-forming, ± succulent, 70–150 cm, glabrous. |
Leaves | opposite, decussate; petiole absent; blade linear to oblong-lanceolate, 30–120 × 3–25 mm, base acute, rounded, cordate or clasping, margins entire, apex acute or subobtuse, sometimes mucronate, surfaces glabrous, abaxial ± glaucous; venation pinnate, midvein prominent. |
opposite; stipules usually distinct, occasionally connate basally (primarily at distal nodes), usually divided into 2–5 subulate to subulate-filiform segments, occasionally forming narrow deltate segments (primarily at distal nodes), 0.8–1.8 mm, glabrous; petiole 3.1–6.3 mm, glabrous; blade ovate to elliptic-oblong, 5–26 × 4–12 mm, base subsymmetric to symmetric, rounded to cuneate, margins entire, apex usually mucronate, rarely acute or obtuse, surfaces glabrous; pinnately veined. |
Involucre | campanulate, 1.2–2.3 × 1.4–2.5 mm, glabrous; glands 4, elliptic, 0.3–0.6 × 1–1.3 mm; horns divergent, thick, tips rounded, dilated, 0.5–1.4 mm. |
campanulate, 1.1–1.8 × 1.5–2.5 mm, glabrous; glands 4, yellowish, sessile or short stipitate, circular to oblong, 0.5–0.7 × 0.5–0.8 mm; appendages white to yellowish, ovate to oblong, occasionally rudimentary, (0–)0.8–1.2 × 0.8–1.5 mm, distal margin entire. |
Staminate flowers | 25–30. |
15–25. |
Pistillate flowers | ovary glabrous; styles 0.7–2.2 mm, 2-fid. |
ovary glabrous; styles 0.7–1 mm, 2-fid 1/2 length. |
Capsules | depressed-globose, 9–12 × 12–16 mm, deeply 3-lobed, tardily dehiscent and appearing indehiscent, mesocarp spongy; cocci rounded, smooth, glabrous; columella 4–5.2 mm. |
ovoid, 4.7–5.5(–6) × 3.6–5.1 mm, glabrous; columella 4.3–5.1 mm. |
Seeds | brownish or blackish, oblong, 4.5–6 × 3–4.2 mm, rugose, irregularly reticulate; caruncle substipitate, hat-shaped, 1.6–2 × 1.2–1.5 mm. |
grayish white to reddish brown mottled, bottle-shaped, strongly dorsiventrally compressed and weakly 3-angled in cross section, (2.8–)4.1–5.2 × 1.2–2(–3.4) mm, smooth; carunclelike structure linear, 0.4–0.5 × 0.1–0.2 mm. |
Cyathial | arrangement: terminal pleiochasial branches 2–4, each 1–2 times 2-branched; pleiochasial bracts cordate-lanceolate, shorter and wider than distal leaves; dichasial bracts distinct, ovate-oblong to lanceolate, base subcordate, margins entire, apex acute; axillary cymose branches 0–10. |
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Cyathia | peduncle 0–0.5 mm. |
solitary at distal nodes; peduncle 1.4–3.6 mm. |
Euphorbia lathyris |
Euphorbia carunculata |
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Phenology | Flowering and fruiting winter–fall. | Flowering and fruiting summer–fall. |
Habitat | Roadsides, cultivated fields, stream banks, waste places. | Sand dunes. |
Elevation | 0–1800 m. (0–5900 ft.) | 400–1300 m. (1300–4300 ft.) |
Distribution |
AZ; CA; CT; ID; IL; MA; MD; MT; NC; OH; OR; PA; SC; TN; TX; VA; WA; WV; BC; ON; QC; Europe; Asia; Africa [Introduced in North America; introduced also in Mexico, South America, Australia]
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KS; NM; OK; TX; Mexico (Chihuahua) |
Discussion | Euphorbia lathyris is most likely native to the central and eastern Mediterranean region, but it is widely cultivated and often locally escaped in temperate regions worldwide, as in the flora area. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia carunculata has a highly localized and scattered distribution. The species is restricted to sand dunes and known from only a handful of localities throughout its relatively wide range. The seeds are unique in being bottle-shaped and strongly laterally compressed, and unlike other members of sect. Anisophyllum, there is a minute, linear, carunclelike protuberance at the hilum. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 304. | FNA vol. 12, p. 262. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Galarhoeus lathyris, Tithymalus lathyris | Chamaesyce carunculata |
Name authority | Linnaeus: Sp. Pl. 1: 457. (1753) — (as lathyrus) | Waterfall: Rhodora 50: 63. (1948) |
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