Euphorbia lathyris |
Euphorbia bilobata |
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caper spurge, euphorbe épurge, gopher plant, gopher spurge, mole plant |
black-seed spurge |
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Habit | Herbs, annual or biennial, with taproot. | Herbs, annual, with slender taproot. |
Stems | erect, unbranched or branched, 200 cm, glabrous, glaucous. |
erect, branched, 10–35 cm, glabrous or strigillose (especially when young and around nodes). |
Leaves | opposite, decussate; petiole absent; blade linear to oblong-lanceolate, 30–120 × 3–25 mm, base acute, rounded, cordate or clasping, margins entire, apex acute or subobtuse, sometimes mucronate, surfaces glabrous, abaxial ± glaucous; venation pinnate, midvein prominent. |
opposite proximally, alternate distally; stipules 0.1–0.2 mm; petiole 1–4(–6) mm, glabrous, sericeous or strigillose; blade linear to narrowly elliptic, 8–52 × 2–7 mm, base attenuate, margins entire, ciliate-strigose, apex acute, abaxial surface sparsely strigillose to sericeous, adaxial surface usually glabrous; venation obscure, only midvein conspicuous. |
Involucre | campanulate, 1.2–2.3 × 1.4–2.5 mm, glabrous; glands 4, elliptic, 0.3–0.6 × 1–1.3 mm; horns divergent, thick, tips rounded, dilated, 0.5–1.4 mm. |
obconic, 0.9–1.5 × 0.9–1.3 mm, strigillose to pilose; glands 5, yellow or pink, U-shaped, 0.2–0.3 × 0.4–0.5 mm; appendages greenish, white, or pink, forming narrow rim around gland, or ovate, oblong, or obovate and usually 2-fid, rarely rudimentary, 0.2–1.4 × 0.2–0.6 mm, entire. |
Staminate flowers | 25–30. |
20–25. |
Pistillate flowers | ovary glabrous; styles 0.7–2.2 mm, 2-fid. |
ovary glabrous, puberulent, strigillose, or pilose; styles 0.5–0.8 mm, 2-fid 1/3–1/2 length. |
Capsules | depressed-globose, 9–12 × 12–16 mm, deeply 3-lobed, tardily dehiscent and appearing indehiscent, mesocarp spongy; cocci rounded, smooth, glabrous; columella 4–5.2 mm. |
oblate, 1.5–2.6 × 2.1–3.3 mm, glabrous or puberulent, strigillose, or pilose; columella 1.2–2.1 mm. |
Seeds | brownish or blackish, oblong, 4.5–6 × 3–4.2 mm, rugose, irregularly reticulate; caruncle substipitate, hat-shaped, 1.6–2 × 1.2–1.5 mm. |
brown to grayish black, narrowly ovoid, 3- or 4-angled in cross section, sometimes obscurely so, 1.3–1.9 × 1–1.4 mm, tuberculate, often with shallow depressions; caruncle absent. |
Cyathial | arrangement: terminal pleiochasial branches 2–4, each 1–2 times 2-branched; pleiochasial bracts cordate-lanceolate, shorter and wider than distal leaves; dichasial bracts distinct, ovate-oblong to lanceolate, base subcordate, margins entire, apex acute; axillary cymose branches 0–10. |
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Cyathia | peduncle 0–0.5 mm. |
solitary at distal nodes or in weakly defined cymes or dichasia, dichasial bracts and distal stem leaves wholly green; peduncle 0.5–3.6 mm, strigillose. |
2n | = 32. |
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Euphorbia lathyris |
Euphorbia bilobata |
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Phenology | Flowering and fruiting winter–fall. | Flowering and fruiting spring–fall. |
Habitat | Roadsides, cultivated fields, stream banks, waste places. | Sandy and rocky soils on slopes and canyon bottoms in pine-juniper woodlands, oak woodlands, grasslands. |
Elevation | 0–1800 m. (0–5900 ft.) | 1400–2600 m. (4600–8500 ft.) |
Distribution |
AZ; CA; CT; ID; IL; MA; MD; MT; NC; OH; OR; PA; SC; TN; TX; VA; WA; WV; BC; ON; QC; Europe; Asia; Africa [Introduced in North America; introduced also in Mexico, South America, Australia]
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AZ; NM; TX; Mexico (Chihuahua, Durango, Sonora)
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Discussion | Euphorbia lathyris is most likely native to the central and eastern Mediterranean region, but it is widely cultivated and often locally escaped in temperate regions worldwide, as in the flora area. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
In Texas, Euphorbia bilobata is known only from Jeff Davis County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 304. | FNA vol. 12, p. 243. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Alectoroctonum |
Sibling taxa | ||
Synonyms | Galarhoeus lathyris, Tithymalus lathyris | |
Name authority | Linnaeus: Sp. Pl. 1: 457. (1753) — (as lathyrus) | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 190. (1859) |
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