Euphorbia lathyris |
Euphorbia mesembrianthemifolia |
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caper spurge, euphorbe épurge, gopher plant, gopher spurge, mole plant |
coastal beach sandmat |
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Habit | Herbs, annual or biennial, with taproot. | Subshrubs or shrubs, perennial, with thickened and often woody rootstock. |
Stems | erect, unbranched or branched, 200 cm, glabrous, glaucous. |
erect to ascending, or nearly decumbent in shifting sand, 25–60 cm, glabrous. |
Leaves | opposite, decussate; petiole absent; blade linear to oblong-lanceolate, 30–120 × 3–25 mm, base acute, rounded, cordate or clasping, margins entire, apex acute or subobtuse, sometimes mucronate, surfaces glabrous, abaxial ± glaucous; venation pinnate, midvein prominent. |
opposite; stipules connate, forming conspicuous, ligulate or deltate scale, short cleft or fringed, 1–1.8 mm, glabrous; petiole 0.5–1 mm, glabrous; blade ovate to elliptic often folded along midrib, 5–12 × 3–8 mm, ± fleshy, base slightly asymmetric, truncate to cordate, partially obscuring stem, margins entire, apex usually obtuse, rarely acute, surfaces yellowish to dark green, glabrous glaucous; obscurely 3–5-veined at base, pinnate distally, only midvein conspicuous. |
Involucre | campanulate, 1.2–2.3 × 1.4–2.5 mm, glabrous; glands 4, elliptic, 0.3–0.6 × 1–1.3 mm; horns divergent, thick, tips rounded, dilated, 0.5–1.4 mm. |
campanulate, 1–1.6 × 0.7–1.2 mm, glabrous; glands 4, brown, usually elliptic, occasionally almost round, 0.2–0.4 × 0.5–0.7 mm, fleshy; appendages white, oblong, rarely rudimentary, 0.2–0.4 × 0.5–0.9 mm, distal margin entire or undulate. |
Staminate flowers | 25–30. |
12–20. |
Pistillate flowers | ovary glabrous; styles 0.7–2.2 mm, 2-fid. |
ovary glabrous; styles 0.3–0.4 mm, 2-fid 1/2 length. |
Capsules | depressed-globose, 9–12 × 12–16 mm, deeply 3-lobed, tardily dehiscent and appearing indehiscent, mesocarp spongy; cocci rounded, smooth, glabrous; columella 4–5.2 mm. |
subglobose subtended by calyxlike structure, 1.5–2 × 2.2–2.8 mm, glabrous; columella 1–1.5 mm. |
Seeds | brownish or blackish, oblong, 4.5–6 × 3–4.2 mm, rugose, irregularly reticulate; caruncle substipitate, hat-shaped, 1.6–2 × 1.2–1.5 mm. |
ashen, broadly ovoid, angled in cross section, faces plump, convex, 1.2–1.3 × 0.9–1.2 mm, obscurely pitted. |
Cyathial | arrangement: terminal pleiochasial branches 2–4, each 1–2 times 2-branched; pleiochasial bracts cordate-lanceolate, shorter and wider than distal leaves; dichasial bracts distinct, ovate-oblong to lanceolate, base subcordate, margins entire, apex acute; axillary cymose branches 0–10. |
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Cyathia | peduncle 0–0.5 mm. |
solitary at distal nodes; peduncle 0.5–1 mm. |
Euphorbia lathyris |
Euphorbia mesembrianthemifolia |
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Phenology | Flowering and fruiting winter–fall. | Flowering and fruiting year-round. |
Habitat | Roadsides, cultivated fields, stream banks, waste places. | Sandy and rocky shores, associated beach scrub. |
Elevation | 0–1800 m. (0–5900 ft.) | 0–10 m. (0–0 ft.) |
Distribution |
AZ; CA; CT; ID; IL; MA; MD; MT; NC; OH; OR; PA; SC; TN; TX; VA; WA; WV; BC; ON; QC; Europe; Asia; Africa [Introduced in North America; introduced also in Mexico, South America, Australia]
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FL; Mexico; West Indies; Bermuda; South America (Colombia, Venezuela) |
Discussion | Euphorbia lathyris is most likely native to the central and eastern Mediterranean region, but it is widely cultivated and often locally escaped in temperate regions worldwide, as in the flora area. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia mesembrianthemifolia is found in the flora area along the sandy and rocky shores of southern Florida from Pinellas and Volusia counties southward. It is one of the few members of subg. Chamaesyce in the flora area that is a shrub or subshrub. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 304. | FNA vol. 12, p. 277. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Galarhoeus lathyris, Tithymalus lathyris | Chamaesyce buxifolia, C. mesembrianthemifolia, E. buxifolia |
Name authority | Linnaeus: Sp. Pl. 1: 457. (1753) — (as lathyrus) | Jacquin: Enum. Syst. Pl., 22. (1760) |
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