Euphorbia ipecacuanhae |
Euphorbia hyssopifolia |
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American ipecac, Carolina ipecac, ipecac spurge |
hyssopleaf sandmat |
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Habit | Herbs, perennial, with deep, stout rootstock. | Herbs, annual, with taproot. |
Stems | decumbent or slightly ascending, branched, often densely clumped, previous year's dead stems not persistent, 17–27 cm, usually glabrous, rarely sparsely villous. |
erect to ascending, 80 cm, sparsely to densely pilose or pilose-crinkled proximally, usually glabrous distally. |
Leaves | alternate; stipules 0.1–0.2 mm; petiole (0–)1–2 mm, glabrous; blade usually linear, obovate, or oblanceolate to orbiculate, rarely filiform, proximal greatly reduced, scalelike, 15–70 × 1.5–13 mm gradually smaller proximally, base cuneate, margins entire, apex rounded, broadly acute, or emarginate, surfaces glabrous; venation occasionally obscure on smaller leaves, midvein conspicuous. |
opposite; stipules usually connate, irregularly lacerate, 0.5–1 mm, usually glabrous, occasionally with few marginal hairs; petiole 1–2 mm, glabrous; blade lanceolate to oblong or falcate, 8–35 × 7–15 mm, base asymmetric, rounded, margins serrulate, apex broadly acute, abaxial surface glabrous or sparsely pilose toward base, adaxial surface glabrous; palmately veined at base, pinnate distally. |
Involucre | yellow or yellow-green, hemispheric, 1–1.2 × 2–2.4 mm, glabrous; glands 5, yellow or yellow-green, obovate or elliptic, 0.7–0.8 × 1–1.2 mm; appendages white or green, often forming narrow rim around distal margin of gland, 0–0.2 mm, entire. |
obconic, 0.9–1.1 × 0.7–0.9 mm, glabrous; glands 4(–5) (5th gland without appendage), yellow-green to maroon, elliptic to circular, 0.1–0.2 × 0.1–0.3 mm; appendages spreading, usually white or turning reddish with age, short reniform or semilunate, 0.1–0.3 × 0.2–0.6 mm, distal margin entire or slightly undulate to crenate. |
Staminate flowers | 10–20. |
4–15. |
Pistillate flowers | ovary glabrous; styles 0.4–0.8 mm, 2-fid at apex. |
ovary glabrous; styles 0.5–0.9 mm, 2-fid 1/2 length. |
Capsules | globose, 2.3–3.4 × 3.5–4.2 mm, glabrous; columella 3–3.1 mm. |
depressed-ovoid, 1.5–1.6 × 1.7–1.8 mm, glabrous; columella 1.5–2 mm. |
Seeds | white or brown, ovoid, 2.3–2.5 × 1.5 mm, angular, with 5 longitudinal ridges, shallowly and irregularly pitted; caruncle absent. |
brown to grayish white, ovoid, slightly 4-angled in cross section, abaxial faces convex, adaxial faces slightly concave to slightly convex, 1–1.4 × 0.7–1.1 mm, with 2–3 prominent transverse ridges that do not interrupt adaxial keel, or coarsely and inconspicuously pitted-reticulate. |
Cyathia | usually in terminal dichasia, sometimes pleiochasia; peduncle 10–50(–70) mm, glabrous. |
solitary or in small, cymose clusters occasionally with bractlike leaves at distal nodes or on congested, axillary branches; peduncle 0.5–2.5 mm. |
2n | = 12, 14. |
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Euphorbia ipecacuanhae |
Euphorbia hyssopifolia |
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Phenology | Flowering and fruiting early spring–early summer. | Flowering and fruiting late spring–early fall. |
Habitat | Pine and pine-oak savannas, pine-oak sand hills, turkey oak scrub, open sand habitats. | Disturbed areas, ditches, gardens. |
Elevation | 0–150 m. (0–500 ft.) | 0–1500 m. (0–4900 ft.) |
Distribution |
CT; DC; DE; GA; MD; NC; NJ; NY; PA; SC; VA
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AL; AZ; FL; GA; LA; MS; NM; SC; TX; UT; Mexico; Central America; South America; West Indies [Introduced in tropical Asia, Africa, Australia]
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Discussion | The vegetative stems of Euphorbia ipecacuanhae are often quite short in proportion to the dichasial or pleiochasial branches, thus superficially plants often appear to have mostly opposite leaves. However, careful examination of the base of the plant will reveal alternate leaves. The leaves are extremely variable in both shape and coloration, and the variation can be pronounced within a population or even on a single plant. M. J. Huft (1979) did not recognize infraspecific taxa within E. ipecacuanhae, and his treatment is followed here. This species is found on the Atlantic coastal plain. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia hyssopifolia is native to the New World tropics and is probably also native to parts of the southern United States. However, at least some of the records from the flora area appear to be from adventive plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 247. | FNA vol. 12, p. 272. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Alectoroctonum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | E. arundelana, Tithymalopsis ipecacuanhae | Chamaesyce hyssopifolia, E. jonesii, E. stenomeres |
Name authority | Linnaeus: Sp. Pl. 1: 455. (1753) | Linnaeus: Syst. Nat. ed. 10, 2: 1048. (1759) |
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