Euphorbia ipecacuanhae |
Euphorbia cumulicola |
|
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American ipecac, Carolina ipecac, ipecac spurge |
coastal dune sandmat |
|
Habit | Herbs, perennial, with deep, stout rootstock. | Herbs, usually annual, rarely perennial, with taproot. |
Stems | decumbent or slightly ascending, branched, often densely clumped, previous year's dead stems not persistent, 17–27 cm, usually glabrous, rarely sparsely villous. |
prostrate, 10–20 cm, glabrous. |
Leaves | alternate; stipules 0.1–0.2 mm; petiole (0–)1–2 mm, glabrous; blade usually linear, obovate, or oblanceolate to orbiculate, rarely filiform, proximal greatly reduced, scalelike, 15–70 × 1.5–13 mm gradually smaller proximally, base cuneate, margins entire, apex rounded, broadly acute, or emarginate, surfaces glabrous; venation occasionally obscure on smaller leaves, midvein conspicuous. |
opposite; stipules distinct, linear-subulate, usually divided into 3–7 linear segments, 0.5–1 mm, glabrous; petiole 0.5–1 mm, glabrous; blade narrowly elliptic to lanceolate, 4–8 × 1–2 mm uniform in size, base asymmetric, obtuse to rounded, margins entire, apex obtuse to acute, surfaces green to reddish flushed, glabrous; only midvein conspicuous. |
Involucre | yellow or yellow-green, hemispheric, 1–1.2 × 2–2.4 mm, glabrous; glands 5, yellow or yellow-green, obovate or elliptic, 0.7–0.8 × 1–1.2 mm; appendages white or green, often forming narrow rim around distal margin of gland, 0–0.2 mm, entire. |
campanulate, 0.9–1.1 × 1–1.3 mm, glabrous; glands 4, green to red, slightly stipitate, subcircular, 0.1 × 0.1 mm; appendages white or pink, fringing edge of gland, short-flabellate, often rudimentary, 0.1–0.2 × 0.1 mm, distal margin crenate or entire. |
Staminate flowers | 10–20. |
5–8. |
Pistillate flowers | ovary glabrous; styles 0.4–0.8 mm, 2-fid at apex. |
ovary glabrous; styles 0.2–0.3 mm, 2-fid at apex to nearly 1/2 length. |
Capsules | globose, 2.3–3.4 × 3.5–4.2 mm, glabrous; columella 3–3.1 mm. |
ovoid or subglobose, 1.8–2 × 2–2.3 mm, glabrous; columella 1.2–1.5 mm. |
Seeds | white or brown, ovoid, 2.3–2.5 × 1.5 mm, angular, with 5 longitudinal ridges, shallowly and irregularly pitted; caruncle absent. |
white to gray-brown, ovoid, terete to bluntly subangled in cross section, 1–1.3 × 1 mm, smooth or minutely pitted, with smooth brown line from top to bottom on adaxial side. |
Cyathia | usually in terminal dichasia, sometimes pleiochasia; peduncle 10–50(–70) mm, glabrous. |
solitary at distal nodes; peduncle 1 mm. |
Euphorbia ipecacuanhae |
Euphorbia cumulicola |
|
Phenology | Flowering and fruiting early spring–early summer. | Flowering and fruiting year-round. |
Habitat | Pine and pine-oak savannas, pine-oak sand hills, turkey oak scrub, open sand habitats. | Sandy oak hammocks, open sandy areas behind mangroves, disturbed sandy sites. |
Elevation | 0–150 m. (0–500 ft.) | 0–10 m. (0–0 ft.) |
Distribution |
CT; DC; DE; GA; MD; NC; NJ; NY; PA; SC; VA
|
FL |
Discussion | The vegetative stems of Euphorbia ipecacuanhae are often quite short in proportion to the dichasial or pleiochasial branches, thus superficially plants often appear to have mostly opposite leaves. However, careful examination of the base of the plant will reveal alternate leaves. The leaves are extremely variable in both shape and coloration, and the variation can be pronounced within a population or even on a single plant. M. J. Huft (1979) did not recognize infraspecific taxa within E. ipecacuanhae, and his treatment is followed here. This species is found on the Atlantic coastal plain. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia cumulicola could be confused with E. bombensis, both of which are widespread in Florida, but there the latter occurs on beaches close to the ocean, whereas E. cumulicola is not a beach-inhabiting species but occurs in more protected sandy habitats such as hammocks or stabilized dunes behind mangroves. Also, E. bombensis has fleshier leaves and larger seeds, and is more compact in habit than E. cumulicola, which is a more sprawling and densely branched plant with leaves that are usually purplish tinged along the margins. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 247. | FNA vol. 12, p. 265. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Alectoroctonum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | E. arundelana, Tithymalopsis ipecacuanhae | Chamaesyce cumulicola |
Name authority | Linnaeus: Sp. Pl. 1: 455. (1753) | (Small) Oudejans: Phytologia 67: 45. (1989) |
Web links |