Euphorbia ipecacuanhae |
Euphorbia chamaesula |
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American ipecac, Carolina ipecac, ipecac spurge |
mountain spurge |
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Habit | Herbs, perennial, with deep, stout rootstock. | Herbs, perennial, with thick rootstock. |
Stems | decumbent or slightly ascending, branched, often densely clumped, previous year's dead stems not persistent, 17–27 cm, usually glabrous, rarely sparsely villous. |
erect, branched, 40–90 cm, glabrous. |
Leaves | alternate; stipules 0.1–0.2 mm; petiole (0–)1–2 mm, glabrous; blade usually linear, obovate, or oblanceolate to orbiculate, rarely filiform, proximal greatly reduced, scalelike, 15–70 × 1.5–13 mm gradually smaller proximally, base cuneate, margins entire, apex rounded, broadly acute, or emarginate, surfaces glabrous; venation occasionally obscure on smaller leaves, midvein conspicuous. |
petiole 0.5–1 mm; blade elliptic to oblong, 8–20(–40) × 3–6 mm, base truncate, rounded, or attenuate, margins entire, apex obtuse or acute, sometimes slightly mucronate, surfaces glabrous; venation inconspicuous, only midvein prominent. |
Involucre | yellow or yellow-green, hemispheric, 1–1.2 × 2–2.4 mm, glabrous; glands 5, yellow or yellow-green, obovate or elliptic, 0.7–0.8 × 1–1.2 mm; appendages white or green, often forming narrow rim around distal margin of gland, 0–0.2 mm, entire. |
turbinate or campanulate, 1.5–2.5 × 1.1–1.9 mm, glabrous; glands 4, crescent-shaped to semicircular, 0.5–0.8 × 1–1.8 mm; horns usually convergent, 0.2–0.8 mm. |
Staminate flowers | 10–20. |
8–12. |
Pistillate flowers | ovary glabrous; styles 0.4–0.8 mm, 2-fid at apex. |
ovary glabrous; styles 1–1.8 mm, 2-fid. |
Capsules | globose, 2.3–3.4 × 3.5–4.2 mm, glabrous; columella 3–3.1 mm. |
depressed-ovoid, 4.3–5 × 5–6 mm, 3-lobed; cocci rounded, smooth, glabrous; columella 3.5–4 mm. |
Seeds | white or brown, ovoid, 2.3–2.5 × 1.5 mm, angular, with 5 longitudinal ridges, shallowly and irregularly pitted; caruncle absent. |
gray to dark brown, ovoid-oblong, truncate at both ends, 2.6–3.4 × 2–2.6 mm, shallowly pitted to almost smooth; caruncle conic, 1 × 0.8 mm. |
Cyathia | usually in terminal dichasia, sometimes pleiochasia; peduncle 10–50(–70) mm, glabrous. |
peduncle 1–3 mm. |
Cyathial | arrangement: terminal pleiochasial branches 3–5(–6), each 3–4 times 2-branched; pleiochasial bracts ovate-lanceolate to slightly subpandurate, similar in size or wider than distal leaves; dichasial bracts distinct, lanceolate to broadly ovate, base usually truncate to rounded or subcordate, sometimes attenuate, margins entire or slightly crenulate, apex usually obtuse to acute, occasionally acuminate; axillary cymose branches 2–8. |
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2n | = 26. |
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Euphorbia ipecacuanhae |
Euphorbia chamaesula |
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Phenology | Flowering and fruiting early spring–early summer. | Flowering and fruiting spring–summer. |
Habitat | Pine and pine-oak savannas, pine-oak sand hills, turkey oak scrub, open sand habitats. | Clearings in ponderosa pine forests, montane roadsides, dry streambeds, creek banks, sandy and gravelly soils. |
Elevation | 0–150 m. (0–500 ft.) | 1700–2700 m. (5600–8900 ft.) |
Distribution |
CT; DC; DE; GA; MD; NC; NJ; NY; PA; SC; VA
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AZ; NM; Mexico (Chihuahua, Sonora)
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Discussion | The vegetative stems of Euphorbia ipecacuanhae are often quite short in proportion to the dichasial or pleiochasial branches, thus superficially plants often appear to have mostly opposite leaves. However, careful examination of the base of the plant will reveal alternate leaves. The leaves are extremely variable in both shape and coloration, and the variation can be pronounced within a population or even on a single plant. M. J. Huft (1979) did not recognize infraspecific taxa within E. ipecacuanhae, and his treatment is followed here. This species is found on the Atlantic coastal plain. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia chamaesula is easily distinguished from other perennial members of subg. Esula in western North America by its larger capsules and the vegetative shoots (without cyathia) that arise from the distal nodes of the stem proximal to the pleiochasia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 247. | FNA vol. 12, p. 300. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Alectoroctonum | Euphorbiaceae > Euphorbia > subg. Esula |
Sibling taxa | ||
Synonyms | E. arundelana, Tithymalopsis ipecacuanhae | Tithymalus chamaesula |
Name authority | Linnaeus: Sp. Pl. 1: 455. (1753) | Boissier: Cent. Euphorb., 38. (1860) |
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