Euphorbia ipecacuanhae |
Euphorbia carunculata |
|
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American ipecac, Carolina ipecac, ipecac spurge |
sand-dune sandmat or spurge |
|
Habit | Herbs, perennial, with deep, stout rootstock. | Herbs, annual, with taproot. |
Stems | decumbent or slightly ascending, branched, often densely clumped, previous year's dead stems not persistent, 17–27 cm, usually glabrous, rarely sparsely villous. |
prostrate, spreading and lanky or occasionally mat-forming, ± succulent, 70–150 cm, glabrous. |
Leaves | alternate; stipules 0.1–0.2 mm; petiole (0–)1–2 mm, glabrous; blade usually linear, obovate, or oblanceolate to orbiculate, rarely filiform, proximal greatly reduced, scalelike, 15–70 × 1.5–13 mm gradually smaller proximally, base cuneate, margins entire, apex rounded, broadly acute, or emarginate, surfaces glabrous; venation occasionally obscure on smaller leaves, midvein conspicuous. |
opposite; stipules usually distinct, occasionally connate basally (primarily at distal nodes), usually divided into 2–5 subulate to subulate-filiform segments, occasionally forming narrow deltate segments (primarily at distal nodes), 0.8–1.8 mm, glabrous; petiole 3.1–6.3 mm, glabrous; blade ovate to elliptic-oblong, 5–26 × 4–12 mm, base subsymmetric to symmetric, rounded to cuneate, margins entire, apex usually mucronate, rarely acute or obtuse, surfaces glabrous; pinnately veined. |
Involucre | yellow or yellow-green, hemispheric, 1–1.2 × 2–2.4 mm, glabrous; glands 5, yellow or yellow-green, obovate or elliptic, 0.7–0.8 × 1–1.2 mm; appendages white or green, often forming narrow rim around distal margin of gland, 0–0.2 mm, entire. |
campanulate, 1.1–1.8 × 1.5–2.5 mm, glabrous; glands 4, yellowish, sessile or short stipitate, circular to oblong, 0.5–0.7 × 0.5–0.8 mm; appendages white to yellowish, ovate to oblong, occasionally rudimentary, (0–)0.8–1.2 × 0.8–1.5 mm, distal margin entire. |
Staminate flowers | 10–20. |
15–25. |
Pistillate flowers | ovary glabrous; styles 0.4–0.8 mm, 2-fid at apex. |
ovary glabrous; styles 0.7–1 mm, 2-fid 1/2 length. |
Capsules | globose, 2.3–3.4 × 3.5–4.2 mm, glabrous; columella 3–3.1 mm. |
ovoid, 4.7–5.5(–6) × 3.6–5.1 mm, glabrous; columella 4.3–5.1 mm. |
Seeds | white or brown, ovoid, 2.3–2.5 × 1.5 mm, angular, with 5 longitudinal ridges, shallowly and irregularly pitted; caruncle absent. |
grayish white to reddish brown mottled, bottle-shaped, strongly dorsiventrally compressed and weakly 3-angled in cross section, (2.8–)4.1–5.2 × 1.2–2(–3.4) mm, smooth; carunclelike structure linear, 0.4–0.5 × 0.1–0.2 mm. |
Cyathia | usually in terminal dichasia, sometimes pleiochasia; peduncle 10–50(–70) mm, glabrous. |
solitary at distal nodes; peduncle 1.4–3.6 mm. |
Euphorbia ipecacuanhae |
Euphorbia carunculata |
|
Phenology | Flowering and fruiting early spring–early summer. | Flowering and fruiting summer–fall. |
Habitat | Pine and pine-oak savannas, pine-oak sand hills, turkey oak scrub, open sand habitats. | Sand dunes. |
Elevation | 0–150 m. (0–500 ft.) | 400–1300 m. (1300–4300 ft.) |
Distribution |
CT; DC; DE; GA; MD; NC; NJ; NY; PA; SC; VA
|
KS; NM; OK; TX; Mexico (Chihuahua) |
Discussion | The vegetative stems of Euphorbia ipecacuanhae are often quite short in proportion to the dichasial or pleiochasial branches, thus superficially plants often appear to have mostly opposite leaves. However, careful examination of the base of the plant will reveal alternate leaves. The leaves are extremely variable in both shape and coloration, and the variation can be pronounced within a population or even on a single plant. M. J. Huft (1979) did not recognize infraspecific taxa within E. ipecacuanhae, and his treatment is followed here. This species is found on the Atlantic coastal plain. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia carunculata has a highly localized and scattered distribution. The species is restricted to sand dunes and known from only a handful of localities throughout its relatively wide range. The seeds are unique in being bottle-shaped and strongly laterally compressed, and unlike other members of sect. Anisophyllum, there is a minute, linear, carunclelike protuberance at the hilum. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 247. | FNA vol. 12, p. 262. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Alectoroctonum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | E. arundelana, Tithymalopsis ipecacuanhae | Chamaesyce carunculata |
Name authority | Linnaeus: Sp. Pl. 1: 455. (1753) | Waterfall: Rhodora 50: 63. (1948) |
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