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American ipecac, Carolina ipecac, ipecac spurge
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Dixie sandmat, southern seaside spurge
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| Herbs, perennial, with deep, stout rootstock. |
Herbs, usually annual, rarely perennial, with taproot. |
decumbent or slightly ascending, branched, often densely clumped, previous year's dead stems not persistent, 17–27 cm, usually glabrous, rarely sparsely villous. |
prostrate or slightly ascending, 10–40 cm, glabrous. |
alternate; stipules 0.1–0.2 mm; petiole (0–)1–2 mm, glabrous; blade usually linear, obovate, or oblanceolate to orbiculate, rarely filiform, proximal greatly reduced, scalelike, 15–70 × 1.5–13 mm gradually smaller proximally, base cuneate, margins entire, apex rounded, broadly acute, or emarginate, surfaces glabrous; venation occasionally obscure on smaller leaves, midvein conspicuous. |
opposite; stipules distinct, linear-subulate, usually divided into 3 linear segments, 1–2 mm, glabrous; petiole 1–2 mm, glabrous; blade oblong or elliptic-oblong, 4–15 × 2–3 mm, base asymmetric to nearly symmetric, obtuse, margins entire, apex acute to mucronate, surfaces green to reddish flushed, glabrous; only midvein conspicuous. |
yellow or yellow-green, hemispheric, 1–1.2 × 2–2.4 mm, glabrous; glands 5, yellow or yellow-green, obovate or elliptic, 0.7–0.8 × 1–1.2 mm; appendages white or green, often forming narrow rim around distal margin of gland, 0–0.2 mm, entire. |
obconic-campanulate, 1.2–1.6 × 1.5–1.7 mm, glabrous; glands 4, green to red, slightly concave, elliptic, oblong, or subcircular, 0.3–0.5 × 0.4–0.6 mm; appendages white or pink, semilunate, fringing edge of gland, sometimes rudimentary, (0–)0.1–0.5(–0.7) × 0.5–0.8 mm, distal margin crenate to entire. |
10–20. |
5–16. |
ovary glabrous; styles 0.4–0.8 mm, 2-fid at apex. |
ovary glabrous; styles 0.2–0.3 mm, 2-fid 1/2 length. |
globose, 2.3–3.4 × 3.5–4.2 mm, glabrous; columella 3–3.1 mm. |
broadly ovoid, 2–2.1 × 2.3–2.5 mm, glabrous; columella 1.5–2 mm. |
white or brown, ovoid, 2.3–2.5 × 1.5 mm, angular, with 5 longitudinal ridges, shallowly and irregularly pitted; caruncle absent. |
ashy white, plumply ovoid, terete to bluntly subangled in cross section, 1.5–1.9 × 1–1.2 mm, smooth or minutely pitted, with smooth brown line from top to bottom on adaxial side. |
usually in terminal dichasia, sometimes pleiochasia; peduncle 10–50(–70) mm, glabrous. |
solitary or in small, cymose clusters at distal nodes; peduncle 0.5–3 mm. |
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| Flowering and fruiting early spring–early summer. |
Flowering and fruiting spring–fall. |
| Pine and pine-oak savannas, pine-oak sand hills, turkey oak scrub, open sand habitats. |
Coastal dunes and sandy habitats. |
| 0–150 m. (0–500 ft.) |
0–30 m. (0–100 ft.) |
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CT; DC; DE; GA; MD; NC; NJ; NY; PA; SC; VA
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AL; FL; GA; LA; MS; NC; SC; TX; VA; Mexico; Central America; n South America (Venezuela)
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The vegetative stems of Euphorbia ipecacuanhae are often quite short in proportion to the dichasial or pleiochasial branches, thus superficially plants often appear to have mostly opposite leaves. However, careful examination of the base of the plant will reveal alternate leaves. The leaves are extremely variable in both shape and coloration, and the variation can be pronounced within a population or even on a single plant. M. J. Huft (1979) did not recognize infraspecific taxa within E. ipecacuanhae, and his treatment is followed here. This species is found on the Atlantic coastal plain. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia bombensis is similar and closely related to E. cumulicola, E. geyeri, and E. polygonifolia. It differs notably from E. polygonifolia in its smaller, plumply ovoid seeds. L. C. Wheeler (1941) suggested that where E. bombensis and E. polygonifolia are sympatric, E. bombensis grows farther away from the shore. Euphorbia bombensis is usually distinguished from E. cumulicola by the latter's smaller, isomorphic leaves that lack any fleshiness, smaller seeds, and diffuse growth habit. Euphorbia bombensis differs from E. geyeri in its usually shorter, less conspicuous involucral gland appendages and its geographic restriction to the coastal plain. However, Wheeler pointed out that plants of E. bombensis from Texas have more or less conspicuous involucral gland appendages. Examination of specimens confirmed that E. bombensis occasionally has conspicuous involucral gland appendages, and because of this, E. bombensis and E. geyeri are difficult to distinguish in Texas. This clade of closely related, sand and dune specialists requires further study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| FNA vol. 12, p. 247. |
FNA vol. 12, p. 261. |
| Euphorbiaceae > Euphorbia > sect. Alectoroctonum |
Euphorbiaceae > Euphorbia > sect. Anisophyllum |
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana |
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana |
| E. arundelana, Tithymalopsis ipecacuanhae |
Chamaesyce ammannioides, C. bombensis, C. ingallsii, E. ammannioides |
| Linnaeus: Sp. Pl. 1: 455. (1753) |
Jacquin: Enum. Syst. Pl., 22. (1760) |
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