Euphorbia ipecacuanhae |
Euphorbia astyla |
|
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American ipecac, Carolina ipecac, ipecac spurge |
alkali spurge, Pecos spurge |
|
Habit | Herbs, perennial, with deep, stout rootstock. | Herbs, perennial, with woody or fibrous-fleshy taproot, 5–12 mm thick. |
Stems | decumbent or slightly ascending, branched, often densely clumped, previous year's dead stems not persistent, 17–27 cm, usually glabrous, rarely sparsely villous. |
decumbent, ascending, or erect, few to many emerging from woody crown, 5–25(–50) cm, glabrous. |
Leaves | alternate; stipules 0.1–0.2 mm; petiole (0–)1–2 mm, glabrous; blade usually linear, obovate, or oblanceolate to orbiculate, rarely filiform, proximal greatly reduced, scalelike, 15–70 × 1.5–13 mm gradually smaller proximally, base cuneate, margins entire, apex rounded, broadly acute, or emarginate, surfaces glabrous; venation occasionally obscure on smaller leaves, midvein conspicuous. |
opposite; stipules connate into deltate scale, 0.2–0.5 mm, minutely lacerate at apex, glabrous; petiole 0–0.2(–0.3) mm, glabrous; blade orbiculate-reniform to acute-cordate, 2–5(–8) × 2–5(–6) mm, base ± asymmetric, cordate to auriculate, sometimes clasping stem, margins entire, apex narrowly acute, surfaces glabrous; 2- or 3-veined from base, but usually only midvein conspicuous. |
Involucre | yellow or yellow-green, hemispheric, 1–1.2 × 2–2.4 mm, glabrous; glands 5, yellow or yellow-green, obovate or elliptic, 0.7–0.8 × 1–1.2 mm; appendages white or green, often forming narrow rim around distal margin of gland, 0–0.2 mm, entire. |
broadly campanulate, 0.8–1.4 × 0.9–1.4 mm, glabrous; glands 4, yellow-green to brownish, oblong, 0.2–0.3 × 0.5–0.7 mm; appendages white, flabellate to oblong, 0.1–0.2(–0.5) × 0.4–0.8 mm, distal margin entire or dentate-crenate. |
Staminate flowers | 10–20. |
22–26. |
Pistillate flowers | ovary glabrous; styles 0.4–0.8 mm, 2-fid at apex. |
ovary glabrous; styles 0.3–0.4 mm, unbranched, thickened-clavate. |
Capsules | globose, 2.3–3.4 × 3.5–4.2 mm, glabrous; columella 3–3.1 mm. |
ovoid and broadly triangular, 1.5–1.9(–2.5) × 1.4–1.6(–2.2) mm, glabrous; columella 1.2–1.8 mm. |
Seeds | white or brown, ovoid, 2.3–2.5 × 1.5 mm, angular, with 5 longitudinal ridges, shallowly and irregularly pitted; caruncle absent. |
white, oblong, 4-angled in cross section, adaxial faces slightly concave, with long raphe between, 1.5–1.8 × 0.7–1 mm, markedly foveolate, with irregular to ± parallel or anastomosing ridges. |
Cyathia | usually in terminal dichasia, sometimes pleiochasia; peduncle 10–50(–70) mm, glabrous. |
solitary at distal nodes; peduncle 0.3–1(–1.5) mm. |
2n | = 28. |
|
Euphorbia ipecacuanhae |
Euphorbia astyla |
|
Phenology | Flowering and fruiting early spring–early summer. | Flowering and fruiting late spring–early fall. |
Habitat | Pine and pine-oak savannas, pine-oak sand hills, turkey oak scrub, open sand habitats. | Desert, grasslands, limestone substrates, usually on very saline or alkaline soils. |
Elevation | 0–150 m. (0–500 ft.) | 700–1100 m. (2300–3600 ft.) |
Distribution |
CT; DC; DE; GA; MD; NC; NJ; NY; PA; SC; VA
|
TX; Mexico (Coahuila, Nuevo León) |
Discussion | The vegetative stems of Euphorbia ipecacuanhae are often quite short in proportion to the dichasial or pleiochasial branches, thus superficially plants often appear to have mostly opposite leaves. However, careful examination of the base of the plant will reveal alternate leaves. The leaves are extremely variable in both shape and coloration, and the variation can be pronounced within a population or even on a single plant. M. J. Huft (1979) did not recognize infraspecific taxa within E. ipecacuanhae, and his treatment is followed here. This species is found on the Atlantic coastal plain. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia astyla is a specialist on halophytic, alkaline soils and is known in the flora area only in part of Pecos County. The species is closely related to E. jejuna but differs in its sessile or sub-sessile leaves with a cordate-auriculate base and involucral gland appendages that are not deeply lobed or cleft. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 247. | FNA vol. 12, p. 261. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Alectoroctonum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | E. arundelana, Tithymalopsis ipecacuanhae | Chamaesyce astyla |
Name authority | Linnaeus: Sp. Pl. 1: 455. (1753) | Engelmann ex Boissier: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 15(2): 40. (1862) |
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