Euphorbia indivisa |
Euphorbia vallis-mortae |
|
---|---|---|
royal sandmat |
Death Valley sandmat |
|
Habit | Herbs, annual or short-lived perennial, with slender taproot to thickened and woody rootstock. | Herbs, perennial, with thickened, woody taproot. |
Stems | prostrate, usually mat-forming, terete to slightly flattened, 40 cm, lower surface glabrous, upper surface strigillose, pilose or villous. |
prostrate to ascending, often mat-forming, 10–45 cm, pilose to villous. |
Leaves | opposite; stipules distinct, entire or divided into 3–4 subulate to filiform segments, 0.8–2 mm, usually pilose, rarely glabrous; petiole 0.5–1 mm, pilose to villous; blade oblong, ovate or narrowly obovate, 3–10(–12) × 2–6 mm, base strongly asymmetric, hemicordate, margins serrulate, apex obtuse to subacute, surfaces glabrous or slightly pilose; 3-veined from base, often only midvein conspicuous. |
opposite; stipules distinct or connate, subulate to filiform, 0.4–1.1 mm, densely tomentose; petiole 0.4–1 mm, pilose to villous; blade suborbiculate to oblong-ovate, 3–8 × 2–6 mm, base slightly asymmetric, rounded, margins entire, apex rounded to obtuse, surfaces pilose to villous; 3-veined at base, midvein conspicuous, venation often obscured by pubescence. |
Involucre | narrowly turbinate, 1–1.2 × 0.4–0.7 mm, pilose; glands 4, yellow to pink, unequal, proximal pair oblong or linear, 0.1 × 0.3–0.4(–0.6) mm, distal pair oblong or subcircular, 0.1 × 0.1–0.2 mm; appendages pink to reddish, unequal, on proximal glands oblique, 0.4–0.8(–1) × 0.8–1.4(–2) mm, on distal glands symmetric, 0.2–0.5 × 0.2–0.5 mm, slightly undulate to slightly crenate. |
obconic-campanulate, 1.2–2.3 × 1–1.8 mm, densely pilose to villous; glands 4, yellow to red, subcircular to oblong, 0.2–0.5 × 0.4–0.9 mm; appendages white, flabellate to oblong, 0.1–0.7 × 0.5–1.9 mm, distal margin entire or crenulate, adaxial surface ciliate-puberulent. |
Staminate flowers | 5–15. |
15–22(–50). |
Pistillate flowers | ovary pilose to strigillose in parts, glabrous in other parts; styles 0.8–1.3 mm, usually unbranched, rarely 2-fid at apex, filiform. |
ovary densely pilose; styles 0.4–0.8 mm, 2-fid 1/2 length. |
Capsules | ovoid-triangular, 1.2–1.5 × 1–1.4 mm, pilose to strigillose in parts, glabrous in other parts; columella 1–1.3 mm. |
ovoid, 1.5–2.2 × 1.8–2.2 mm, tomentose; columella 1.2–1.6 mm. |
Seeds | brown to light gray, ovoid, 4-angled in cross section, 0.8–1 × 0.4–0.5 mm, with 4 or 5 deep transverse sulci alternating with low transverse ridges. |
white, gray, or light brown, ovoid, sharply 4-angled in cross section, abaxial faces slightly convex, adaxial faces concave, 1.2–1.7 × 0.6–0.9 mm, smooth. |
Cyathia | usually in small cymose clusters on congested, axillary branches; peduncle rudimentary or to 0.2 mm. |
solitary at distal nodes; peduncle 0.5–1.8 mm. |
Euphorbia indivisa |
Euphorbia vallis-mortae |
|
Phenology | Flowering and fruiting summer–fall. | Flowering and fruiting late spring–fall. |
Habitat | Grasslands, oak forests, oak-mesquite woodlands, oak-juniper communities, rarely entering desert scrub. | Roadsides, desert scrub, streamsides, sandy washes. |
Elevation | 1000–2000 m. (3300–6600 ft.) | 700–2000 m. (2300–6600 ft.) |
Distribution |
AZ; NM; TX; Mexico
|
CA |
Discussion | Euphorbia indivisa is characteristic of grasslands and oak woodlands from extreme western Texas to southeastern Arizona. The species is often treated as a synonym of E. dioeca Kunth, but the two species are readily separable on the basis of their seeds. The seeds of E. indivisa possess deep transverse sulci, whereas those of E. dioeca are merely rippled or with low transverse ridges. Euphorbia dioeca is a weedy species that occurs widely throughout tropical America but has yet to be encountered within the flora area. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The specific epithet of Euphorbia vallis-mortae is a misnomer because the species does not occur in Death Valley; instead, it is found at the transition of the northern edge of the Mojave Desert and the foothills of the southern Sierra Nevada in Inyo, Kern, and San Bernardino counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 273. | FNA vol. 12, p. 291. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | E. dioeca var. indivisa, Chamaesyce indivisa | Chamaesyce vallis-mortae |
Name authority | (Engelmann) Tidestrom: Proc. Biol. Soc. Wash. 48: 40. (1935) | (Millspaugh) J. T. Howell: Madroño 2: 19. (1931) |
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