Euphorbia indivisa |
Euphorbia jejuna |
|
---|---|---|
royal sandmat |
dwarf broomspurge |
|
Habit | Herbs, annual or short-lived perennial, with slender taproot to thickened and woody rootstock. | Herbs, perennial, with woody or fibrous-fleshy, napiform, branched or tuberous taproot, 3–15 mm thick. |
Stems | prostrate, usually mat-forming, terete to slightly flattened, 40 cm, lower surface glabrous, upper surface strigillose, pilose or villous. |
ascending to erect, densely emerging from woody crown, 5–15 cm, glabrous. |
Leaves | opposite; stipules distinct, entire or divided into 3–4 subulate to filiform segments, 0.8–2 mm, usually pilose, rarely glabrous; petiole 0.5–1 mm, pilose to villous; blade oblong, ovate or narrowly obovate, 3–10(–12) × 2–6 mm, base strongly asymmetric, hemicordate, margins serrulate, apex obtuse to subacute, surfaces glabrous or slightly pilose; 3-veined from base, often only midvein conspicuous. |
opposite; stipules connate into lacerate or 2-fid, lanceolate or deltate scale, 0.5–0.9 mm, glabrous; petiole 0.3–0.9 mm, glabrous; blade orbiculate-obovate, ovate, or elliptic, 3–6(–8) × 1.8–5 mm, base moderately asymmetric, rounded to truncate, margins entire, apex blunt to acute, surfaces glabrous; 2–3-veined from base but usually only midvein conspicuous. |
Involucre | narrowly turbinate, 1–1.2 × 0.4–0.7 mm, pilose; glands 4, yellow to pink, unequal, proximal pair oblong or linear, 0.1 × 0.3–0.4(–0.6) mm, distal pair oblong or subcircular, 0.1 × 0.1–0.2 mm; appendages pink to reddish, unequal, on proximal glands oblique, 0.4–0.8(–1) × 0.8–1.4(–2) mm, on distal glands symmetric, 0.2–0.5 × 0.2–0.5 mm, slightly undulate to slightly crenate. |
broadly campanulate, 1.2–1.5 × 0.8–1.2 mm, glabrous; glands 4, yellowish to green or purplish, oblong, 0.2–0.4 × 0.5–1 mm; appendages erect or spreading, white, 0.3–0.6 × 0.8–1.2 mm, usually deeply dissected into 4–5 acuminate lobes, rarely undivided, when divided distal margin rarely entire or crenate. |
Staminate flowers | 5–15. |
12–35. |
Pistillate flowers | ovary pilose to strigillose in parts, glabrous in other parts; styles 0.8–1.3 mm, usually unbranched, rarely 2-fid at apex, filiform. |
ovary glabrous; styles 0.4–0.5 mm, unbranched thickened-clavate. |
Capsules | ovoid-triangular, 1.2–1.5 × 1–1.4 mm, pilose to strigillose in parts, glabrous in other parts; columella 1–1.3 mm. |
ovoid and broadly triangular, 1.8–2.2(–2.7) × 1.5–2.1 mm, glabrous; columella 1.5–2.2 mm. |
Seeds | brown to light gray, ovoid, 4-angled in cross section, 0.8–1 × 0.4–0.5 mm, with 4 or 5 deep transverse sulci alternating with low transverse ridges. |
whitish, oblong, 4-angled in cross section, adaxial faces concave, with long raphe between, 1.5–2(–2.3) × 0.6–0.8 mm, dimpled with faint irregular transverse wrinkles or with up to 10 low, rounded transverse ridges. |
Cyathia | usually in small cymose clusters on congested, axillary branches; peduncle rudimentary or to 0.2 mm. |
solitary at distal nodes; peduncle 0.8–1.5 mm. |
Euphorbia indivisa |
Euphorbia jejuna |
|
Phenology | Flowering and fruiting summer–fall. | Flowering and fruiting early spring–summer. |
Habitat | Grasslands, oak forests, oak-mesquite woodlands, oak-juniper communities, rarely entering desert scrub. | Thin calcareous soils (caliche) on limestone hills. |
Elevation | 1000–2000 m. (3300–6600 ft.) | 500–900 m. (1600–3000 ft.) |
Distribution |
AZ; NM; TX; Mexico
|
TX; Mexico (Chihuahua, Coahuila) |
Discussion | Euphorbia indivisa is characteristic of grasslands and oak woodlands from extreme western Texas to southeastern Arizona. The species is often treated as a synonym of E. dioeca Kunth, but the two species are readily separable on the basis of their seeds. The seeds of E. indivisa possess deep transverse sulci, whereas those of E. dioeca are merely rippled or with low transverse ridges. Euphorbia dioeca is a weedy species that occurs widely throughout tropical America but has yet to be encountered within the flora area. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia jejuna is known in the flora area from only a few collections in Mitchell, Nolan, Terrell, and Val Verde counties. The species is very similar to E. astyla, but differs in its more deeply divided involucral gland appendages and more definitely petiolate, rounder leaves. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 273. | FNA vol. 12, p. 274. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | E. dioeca var. indivisa, Chamaesyce indivisa | Chamaesyce jejuna |
Name authority | (Engelmann) Tidestrom: Proc. Biol. Soc. Wash. 48: 40. (1935) | M. C. Johnston & Warnock: SouthW. Naturalist 5: 97, fig. [p. 98]. (1960) |
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