Euphorbia indivisa |
Euphorbia chamaesula |
|
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royal sandmat |
mountain spurge |
|
Habit | Herbs, annual or short-lived perennial, with slender taproot to thickened and woody rootstock. | Herbs, perennial, with thick rootstock. |
Stems | prostrate, usually mat-forming, terete to slightly flattened, 40 cm, lower surface glabrous, upper surface strigillose, pilose or villous. |
erect, branched, 40–90 cm, glabrous. |
Leaves | opposite; stipules distinct, entire or divided into 3–4 subulate to filiform segments, 0.8–2 mm, usually pilose, rarely glabrous; petiole 0.5–1 mm, pilose to villous; blade oblong, ovate or narrowly obovate, 3–10(–12) × 2–6 mm, base strongly asymmetric, hemicordate, margins serrulate, apex obtuse to subacute, surfaces glabrous or slightly pilose; 3-veined from base, often only midvein conspicuous. |
petiole 0.5–1 mm; blade elliptic to oblong, 8–20(–40) × 3–6 mm, base truncate, rounded, or attenuate, margins entire, apex obtuse or acute, sometimes slightly mucronate, surfaces glabrous; venation inconspicuous, only midvein prominent. |
Involucre | narrowly turbinate, 1–1.2 × 0.4–0.7 mm, pilose; glands 4, yellow to pink, unequal, proximal pair oblong or linear, 0.1 × 0.3–0.4(–0.6) mm, distal pair oblong or subcircular, 0.1 × 0.1–0.2 mm; appendages pink to reddish, unequal, on proximal glands oblique, 0.4–0.8(–1) × 0.8–1.4(–2) mm, on distal glands symmetric, 0.2–0.5 × 0.2–0.5 mm, slightly undulate to slightly crenate. |
turbinate or campanulate, 1.5–2.5 × 1.1–1.9 mm, glabrous; glands 4, crescent-shaped to semicircular, 0.5–0.8 × 1–1.8 mm; horns usually convergent, 0.2–0.8 mm. |
Staminate flowers | 5–15. |
8–12. |
Pistillate flowers | ovary pilose to strigillose in parts, glabrous in other parts; styles 0.8–1.3 mm, usually unbranched, rarely 2-fid at apex, filiform. |
ovary glabrous; styles 1–1.8 mm, 2-fid. |
Capsules | ovoid-triangular, 1.2–1.5 × 1–1.4 mm, pilose to strigillose in parts, glabrous in other parts; columella 1–1.3 mm. |
depressed-ovoid, 4.3–5 × 5–6 mm, 3-lobed; cocci rounded, smooth, glabrous; columella 3.5–4 mm. |
Seeds | brown to light gray, ovoid, 4-angled in cross section, 0.8–1 × 0.4–0.5 mm, with 4 or 5 deep transverse sulci alternating with low transverse ridges. |
gray to dark brown, ovoid-oblong, truncate at both ends, 2.6–3.4 × 2–2.6 mm, shallowly pitted to almost smooth; caruncle conic, 1 × 0.8 mm. |
Cyathia | usually in small cymose clusters on congested, axillary branches; peduncle rudimentary or to 0.2 mm. |
peduncle 1–3 mm. |
Cyathial | arrangement: terminal pleiochasial branches 3–5(–6), each 3–4 times 2-branched; pleiochasial bracts ovate-lanceolate to slightly subpandurate, similar in size or wider than distal leaves; dichasial bracts distinct, lanceolate to broadly ovate, base usually truncate to rounded or subcordate, sometimes attenuate, margins entire or slightly crenulate, apex usually obtuse to acute, occasionally acuminate; axillary cymose branches 2–8. |
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2n | = 26. |
|
Euphorbia indivisa |
Euphorbia chamaesula |
|
Phenology | Flowering and fruiting summer–fall. | Flowering and fruiting spring–summer. |
Habitat | Grasslands, oak forests, oak-mesquite woodlands, oak-juniper communities, rarely entering desert scrub. | Clearings in ponderosa pine forests, montane roadsides, dry streambeds, creek banks, sandy and gravelly soils. |
Elevation | 1000–2000 m. (3300–6600 ft.) | 1700–2700 m. (5600–8900 ft.) |
Distribution |
AZ; NM; TX; Mexico
|
AZ; NM; Mexico (Chihuahua, Sonora)
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Discussion | Euphorbia indivisa is characteristic of grasslands and oak woodlands from extreme western Texas to southeastern Arizona. The species is often treated as a synonym of E. dioeca Kunth, but the two species are readily separable on the basis of their seeds. The seeds of E. indivisa possess deep transverse sulci, whereas those of E. dioeca are merely rippled or with low transverse ridges. Euphorbia dioeca is a weedy species that occurs widely throughout tropical America but has yet to be encountered within the flora area. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia chamaesula is easily distinguished from other perennial members of subg. Esula in western North America by its larger capsules and the vegetative shoots (without cyathia) that arise from the distal nodes of the stem proximal to the pleiochasia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 273. | FNA vol. 12, p. 300. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > subg. Esula |
Sibling taxa | ||
Synonyms | E. dioeca var. indivisa, Chamaesyce indivisa | Tithymalus chamaesula |
Name authority | (Engelmann) Tidestrom: Proc. Biol. Soc. Wash. 48: 40. (1935) | Boissier: Cent. Euphorb., 38. (1860) |
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