Euphorbia hyssopifolia |
Euphorbia serpens |
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hyssopleaf sandmat |
creeping spurge, Matted sandmat, Matted spurge, round-leafed spurge |
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Habit | Herbs, annual, with taproot. | Herbs, annual, with slender taproot. |
Stems | erect to ascending, 80 cm, sparsely to densely pilose or pilose-crinkled proximally, usually glabrous distally. |
prostrate, frequently mat-forming and rooting at nodes, 15–50 cm, glabrous. |
Leaves | opposite; stipules usually connate, irregularly lacerate, 0.5–1 mm, usually glabrous, occasionally with few marginal hairs; petiole 1–2 mm, glabrous; blade lanceolate to oblong or falcate, 8–35 × 7–15 mm, base asymmetric, rounded, margins serrulate, apex broadly acute, abaxial surface glabrous or sparsely pilose toward base, adaxial surface glabrous; palmately veined at base, pinnate distally. |
opposite; stipules connate into conspicuous, deltate scale, white to pink, membranaceous, 0.5–1(–1.2) mm, glabrous; petiole less than 1 mm, glabrous; blade ovate, oblong or orbiculate, 2–7(–9) × 2–6 mm, base asymmetric, rounded to subcordate, margins entire, apex rounded, surfaces without red blotch, glabrous; usually only midvein conspicuous. |
Involucre | obconic, 0.9–1.1 × 0.7–0.9 mm, glabrous; glands 4(–5) (5th gland without appendage), yellow-green to maroon, elliptic to circular, 0.1–0.2 × 0.1–0.3 mm; appendages spreading, usually white or turning reddish with age, short reniform or semilunate, 0.1–0.3 × 0.2–0.6 mm, distal margin entire or slightly undulate to crenate. |
campanulate to turbinate, 0.3–0.7(–1) × 0.4–0.6 mm, glabrous; glands 4, yellow, oblong, 0.1 × 0.2 mm; appendages white to pinkish, equal, forming narrow rim at edge of gland, 0.1–0.2 × 0.2–0.3 mm, distal margin entire or crenulate. |
Staminate flowers | 4–15. |
5–10. |
Pistillate flowers | ovary glabrous; styles 0.5–0.9 mm, 2-fid 1/2 length. |
ovary glabrous; styles 0.2 mm, 2-fid 1/2 to nearly entire length. |
Capsules | depressed-ovoid, 1.5–1.6 × 1.7–1.8 mm, glabrous; columella 1.5–2 mm. |
broadly ovoid, oblate, or subglobose, 1.3–1.4 × 1.3–1.7 mm, glabrous; columella (0.9–)1–1.2 mm. |
Seeds | brown to grayish white, ovoid, slightly 4-angled in cross section, abaxial faces convex, adaxial faces slightly concave to slightly convex, 1–1.4 × 0.7–1.1 mm, with 2–3 prominent transverse ridges that do not interrupt adaxial keel, or coarsely and inconspicuously pitted-reticulate. |
white to gray or light pink, ovoid, bluntly 3–4-angled in cross section, 0.7–1.1 × 0.4–0.7 mm, smooth. |
Cyathia | solitary or in small, cymose clusters occasionally with bractlike leaves at distal nodes or on congested, axillary branches; peduncle 0.5–2.5 mm. |
solitary at distal nodes; peduncle 0.5–1(–2.5) mm. |
2n | = 12, 14. |
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Euphorbia hyssopifolia |
Euphorbia serpens |
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Phenology | Flowering and fruiting late spring–early fall. | Flowering and fruiting year-round (in warmer areas) or summer (in temperate regions). |
Habitat | Disturbed areas, ditches, gardens. | Mostly sandy or well-drained soils, desert scrub, coastal scrub, chaparral, oak and juniper woodlands, sand dunes, riparian forests, mesquite grasslands, prairies, coniferous and deciduous hardwood forests, disturbed areas. |
Elevation | 0–1500 m. (0–4900 ft.) | 0–2000 m. (0–6600 ft.) |
Distribution |
AL; AZ; FL; GA; LA; MS; NM; SC; TX; UT; Mexico; Central America; South America; West Indies [Introduced in tropical Asia, Africa, Australia]
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AL; AR; AZ; CA; CO; FL; GA; IA; IL; IN; KS; KY; LA; MI; MO; MS; MT; ND; NE; NJ; NM; OH; OK; PA; SD; TN; TX; UT; VT; WY; ON; SK; Mexico; Central America; South America; West Indies; Bermuda [Introduced in Europe, Asia, Africa, Australia]
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Discussion | Euphorbia hyssopifolia is native to the New World tropics and is probably also native to parts of the southern United States. However, at least some of the records from the flora area appear to be from adventive plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia serpens is one of the most widespread species of the genus in the New World. While it may be indigenous to a portion of the flora area, probably in the warmer, southern part of its range, it is weedy and has likely been introduced in many parts of the flora area, such as Canada and the eastern United States. It is also widely distributed in the Old World, where it is certainly introduced. The strictly prostrate habit with stems rooting at the nodes is characteristic. Euphorbia serpens is often confused with E. albomarginata, a species distributed in the southwestern United States; in addition to the features mentioned in the key, an easy and reliable way to distinguish between them is by the size of the involucral gland appendages: those of E. albomarginata are conspicuous to the naked eye, whereas those of E. serpens are inconspicuous. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 272. | FNA vol. 12, p. 286. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Chamaesyce hyssopifolia, E. jonesii, E. stenomeres | Chamaesyce serpens |
Name authority | Linnaeus: Syst. Nat. ed. 10, 2: 1048. (1759) | Kunth: in A. von Humboldt et al., Nov. Gen. Sp. 2(fol.): 41; 2(qto.): 52. (1817) |
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