Euphorbia hyssopifolia |
Euphorbia heterophylla |
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hyssopleaf sandmat |
Mexican fireplant, painted euphorbia |
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Habit | Herbs, annual, with taproot. | Herbs, annual, with taproot. |
Stems | erect to ascending, 80 cm, sparsely to densely pilose or pilose-crinkled proximally, usually glabrous distally. |
erect-ascending, 20–100 cm, sparsely pilose to villous; branches ± straight. |
Leaves | opposite; stipules usually connate, irregularly lacerate, 0.5–1 mm, usually glabrous, occasionally with few marginal hairs; petiole 1–2 mm, glabrous; blade lanceolate to oblong or falcate, 8–35 × 7–15 mm, base asymmetric, rounded, margins serrulate, apex broadly acute, abaxial surface glabrous or sparsely pilose toward base, adaxial surface glabrous; palmately veined at base, pinnate distally. |
usually alternate, occasionally opposite proximally; petiole 10–50 mm, pilose; blade narrowly lanceolate to elliptic or broadly obovate (then usually pandurate and 4-lobed), often polymorphic on single plants, 30–200 × 20–140 mm, base acute, margins sparsely glandular-serrulate, hirtellous, flat, apex acute to obtuse, abaxial surface sparsely appressed-pilose, adaxial surface sparsely pilosulous to glabrate; venation pinnate, midvein prominent. |
Involucre | obconic, 0.9–1.1 × 0.7–0.9 mm, glabrous; glands 4(–5) (5th gland without appendage), yellow-green to maroon, elliptic to circular, 0.1–0.2 × 0.1–0.3 mm; appendages spreading, usually white or turning reddish with age, short reniform or semilunate, 0.1–0.3 × 0.2–0.6 mm, distal margin entire or slightly undulate to crenate. |
usually campanulate, occasionally nearly hemispheric, 1.5–1.9 × 1.2–1.8 mm, glabrous; involucral lobes divided into several linear, smooth lobes; gland 1, yellow-green, stipitate, clavate, 1–1.4 × 1–1.2 mm, opening circular (occasionally flattened from pressing), with annular rim, glabrous; appendages absent. |
Staminate flowers | 4–15. |
8–15. |
Pistillate flowers | ovary glabrous; styles 0.5–0.9 mm, 2-fid 1/2 length. |
ovary glabrous or puberulent; styles 0.8–1.3 mm, 2-fid 1/2 to nearly entire length. |
Capsules | depressed-ovoid, 1.5–1.6 × 1.7–1.8 mm, glabrous; columella 1.5–2 mm. |
broadly ovoid, 2.8–3.8 × 4–5.3 mm, 3-lobed, usually glabrous, rarely sparsely puberulent; columella 2.1–2.8 mm. |
Seeds | brown to grayish white, ovoid, slightly 4-angled in cross section, abaxial faces convex, adaxial faces slightly concave to slightly convex, 1–1.4 × 0.7–1.1 mm, with 2–3 prominent transverse ridges that do not interrupt adaxial keel, or coarsely and inconspicuously pitted-reticulate. |
brown-gray to ashy gray, broadly deltoid, 2.4–2.8 × 1.9–2.4 mm, angular in cross section, dorsal face strongly acute-carinate, tuberculate, with broad rounded tubercles in 2 rows; caruncle 0.1 mm. |
Cyathia | solitary or in small, cymose clusters occasionally with bractlike leaves at distal nodes or on congested, axillary branches; peduncle 0.5–2.5 mm. |
peduncle 0.9–1.5 mm. |
Cyathial | arrangement: terminal dichasial branches usually 2, occasionally reduced to congested cyme, 1–2-branched (often congested and difficult to discern); pleiochasial bracts 2–4, often whorled, wholly green or paler green at base, similar in shape and size to distal leaves; dichasial bracts highly reduced, rarely absent in highly congested clusters. |
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2n | = 12, 14. |
= 28. |
Euphorbia hyssopifolia |
Euphorbia heterophylla |
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Phenology | Flowering and fruiting late spring–early fall. | Flowering and fruiting nearly year-round. |
Habitat | Disturbed areas, ditches, gardens. | Disturbed areas, roadsides. |
Elevation | 0–1500 m. (0–4900 ft.) | 0–500 m. (0–1600 ft.) |
Distribution |
AL; AZ; FL; GA; LA; MS; NM; SC; TX; UT; Mexico; Central America; South America; West Indies [Introduced in tropical Asia, Africa, Australia]
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Mexico; Central America; South America [Introduced, Ala., Ariz., Calif., Fla., Ga., La., Miss., N.Mex., Tex.; introduced also in Eurasia, Africa]
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Discussion | Euphorbia hyssopifolia is native to the New World tropics and is probably also native to parts of the southern United States. However, at least some of the records from the flora area appear to be from adventive plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia heterophylla occurs from the southern United States, where it is likely naturalized, south through Mexico and Central America to South America. Because of its weediness, the precise native range in tropical and subtropical parts of the New World is not well understood. It has become widely established also in warm areas of the Old World. Leaf shape in this species is highly polymorphic within both populations and individuals. Euphorbia heterophylla can appear superficially similar to E. cyathophora but differs in its stipitate, circular involucral glands and its floral bracts that are at most very pale at the base (never colored as is typical in E. cyathophora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 272. | FNA vol. 12, p. 323. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Poinsettia |
Sibling taxa | ||
Synonyms | Chamaesyce hyssopifolia, E. jonesii, E. stenomeres | E. geniculata, Poinsettia geniculata, P. heterophylla |
Name authority | Linnaeus: Syst. Nat. ed. 10, 2: 1048. (1759) | Linnaeus: Sp. Pl. 1: 453. (1753) |
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