Euphorbia hypericifolia |
Euphorbia polycarpa |
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graceful sandmat, graceful spurge |
smallseed sandmat |
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Habit | Herbs, annual, with taproot. | |
Stems | erect to ascending, 15–50 cm, glabrous. |
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Leaves | opposite; stipules connate, deltate, usually entire, sometimes laciniate-fringed at tip, 1.5–2.2 mm, glabrous; petiole 1–3 mm, glabrous; blade obliquely oblong-oblanceolate, 10–35 × 7–15 mm, base asymmetric, oblique, margins serrate or serrulate, especially toward apex, apex broadly acute, surfaces glabrous; palmately veined at base, pinnate distally. |
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Involucre | obconic, 0.9–1.1 × 0.4–0.9 mm, glabrous; glands 4, yellow-green to brown, stipitate, subcircular, 0.2 × 0.2 mm, occasionally nearly rudimentary; appendages absent on smaller glands or white to pink, shape highly variable, usually round to ± elliptic, 0.3–0.4 × 0.5–0.7 mm, distal margin entire. |
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Staminate flowers | (0–)2–20. |
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Pistillate flowers | ovary glabrous; styles 0.4 mm, 2-fid 1/2 length. |
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Capsules | depressed-globoid, 1.3–1.4 × 1.1–1.5 mm, glabrous; columella 1–1.1 mm. |
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Seeds | with very thin whitish mucilaginous coat over light brown testa below, ovoid-triangular, bluntly 4-angled in cross section, 0.9–1.1 × 0.5 mm, with shallow irregular depressions alternating with low, smooth ridges. |
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Cyathia | in dense, axillary and terminal, capitate glomerules with reduced, bractlike leaves subtending cyathia; peduncle 0.5–1.8 mm. |
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Euphorbia | polycarpa is a highly variable species distributed throughout the Baja California peninsula, eastern Sonora, and the arid southwestern United States from southwestern Arizona to southern Nevada and central California. |
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l | . |
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c | . |
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Wheeler | (1941) divided the species into seven varieties, the majority of which occur in Baja California Sur. |
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He | reported two of these for the United States: var. hirtella and var. polycarpa. |
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Variety | hirtella, as the name suggests, was applied to hairy plants. |
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However | , it is here treated as a synonym of var. polycarpa, because variation in pubescence shows no geographic segregation and ranges along a continuum from glabrous or sparsely to densely hairy, and because glabrous and hairy branches can occur on the same individual. |
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Euphorbia hypericifolia |
Euphorbia polycarpa |
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Phenology | Flowering and fruiting early spring–late fall. | |
Habitat | Open, disturbed areas, nurseries. | |
Elevation | 0–200 m. (0–700 ft.) | |
Distribution |
AL; AR; FL; GA; LA; OK; SC; TX; Central America; South America; West Indies [Introduced in Asia, Pacific Islands]
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AZ; CA; NV; nw Mexico
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Discussion | Euphorbia hypericifolia is native to the New World tropics, and it is most likely adventive in the flora area (where it is most widely distributed in Florida and Texas). Reports from Arizona, California, and Maryland likely represent waifs or misidentifications. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 5 (1 in the flora). Some of the varieties occurring in Mexico are highly divergent and appear to be sufficiently distinct to merit recognition as species. However, within the flora area, Euphorbia polycarpa is relatively uniform, and the only noteworthy variation involves the size of the involucral gland appendages. In the portion of the California Floristic Province occupied by E. polycarpa (Los Angeles, Orange, western Riverside, western San Diego, and Ventura counties) the appendages are conspicuously larger than those of plants throughout the remainder of its range in the southwestern United States (0.4–1.2 versus 0.1–0.3 mm). Large-appendaged plants are also common in Baja California and correspond well with the type collection. The taxonomic significance of this variation is not clear at this time, and the small-appendaged plants may merit segregation as an infraspecific taxon. However, no formal changes are proposed, awaiting a comprehensive review of the species throughout its range. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 272. | FNA vol. 12, p. 283. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Subordinate taxa | ||
Synonyms | Chamaesyce glomerifera, C. hypericifolia, E. glomerifera | Chamaesyce polycarpa |
Name authority | Linnaeus: Sp. Pl. 1: 454. (1753) | Bentham: Bot. Voy. Sulphur, 50. (1844) |
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