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graceful sandmat, graceful spurge

red-gland spurge, squaw sandmat

Habit Herbs, annual, with taproot. Herbs, perennial, with moderately to strongly thickened rootstock.
Stems

erect to ascending, 15–50 cm, glabrous.

ascending to erect, 5–20 cm, sericeous to appressed-villous.

Leaves

opposite;

stipules connate, deltate, usually entire, sometimes laciniate-fringed at tip, 1.5–2.2 mm, glabrous;

petiole 1–3 mm, glabrous;

blade obliquely oblong-oblanceolate, 10–35 × 7–15 mm, base asymmetric, oblique, margins serrate or serrulate, especially toward apex, apex broadly acute, surfaces glabrous; palmately veined at base, pinnate distally.

opposite;

stipules distinct (lower side) and connate (upper side), linear, 0.5–1 mm, densely pilose;

petiole 0.8–1.5 mm, tomentose;

blade ovate, 1.2–5 × 0.8–2.9 mm, base asymmetric, hemicordate, margins entire, apex rounded to acute, surfaces tomentose;

venation inconspicuous.

Involucre

obconic, 0.9–1.1 × 0.4–0.9 mm, glabrous;

glands 4, yellow-green to brown, stipitate, subcircular, 0.2 × 0.2 mm, occasionally nearly rudimentary;

appendages absent on smaller glands or white to pink, shape highly variable, usually round to ± elliptic, 0.3–0.4 × 0.5–0.7 mm, distal margin entire.

campanulate, 0.6–1.1 × 0.7–1 mm, tomentose;

glands 4, deep red to purple, elliptic, 0.3–0.4 × 0.4–0.7 mm;

appendages white or becoming pink with age, oblong to flabellate, 0.4–0.7(–1) × 0.7–1.2 mm, distal margin entire or erose.

Staminate flowers

(0–)2–20.

45–80.

Pistillate flowers

ovary glabrous;

styles 0.4 mm, 2-fid 1/2 length.

ovary tomentose, styles 0.5–0.8 mm, 2-fid nearly entire length.

Capsules

depressed-globoid, 1.3–1.4 × 1.1–1.5 mm, glabrous;

columella 1–1.1 mm.

ovoid, 1.4–1.8 × 1.4–1.7 mm, tomentose;

columella 1.2–1.5 mm.

Seeds

with very thin whitish mucilaginous coat over light brown testa below, ovoid-triangular, bluntly 4-angled in cross section, 0.9–1.1 × 0.5 mm, with shallow irregular depressions alternating with low, smooth ridges.

gray to tan, oblong, 4-angled in cross section, 1–1.2 × 0.4–0.6 mm, smooth to wrinkled or alveolate.

Cyathia

in dense, axillary and terminal, capitate glomerules with reduced, bractlike leaves subtending cyathia;

peduncle 0.5–1.8 mm.

solitary at distal nodes;

peduncle (0.6–)1.4–1.9 mm.

Euphorbia hypericifolia

Euphorbia melanadenia

Phenology Flowering and fruiting early spring–late fall. Flowering and fruiting year-round.
Habitat Open, disturbed areas, nurseries. Rocky slopes, river washes, dry to wet soils.
Elevation 0–200 m. (0–700 ft.) 400–1400 m. (1300–4600 ft.)
Distribution
from FNA
AL; AR; FL; GA; LA; OK; SC; TX; Central America; South America; West Indies [Introduced in Asia, Pacific Islands]
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; Mexico (Baja California, Sonora)
[WildflowerSearch map]
[BONAP county map]
Discussion

Euphorbia hypericifolia is native to the New World tropics, and it is most likely adventive in the flora area (where it is most widely distributed in Florida and Texas). Reports from Arizona, California, and Maryland likely represent waifs or misidentifications.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia melanadenia is similar in appearance to E. cinerascens, but E. melanadenia has conspicuous involucral gland appendages whereas E. cinerascens has inconspicuous appendages or lacks them entirely. Euphorbia melanadenia occurs in Arizona and southern California, whereas E. cinerascens is found only in southern and western Texas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 272. FNA vol. 12, p. 276.
Parent taxa Euphorbiaceae > Euphorbia > sect. Anisophyllum Euphorbiaceae > Euphorbia > sect. Anisophyllum
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms Chamaesyce glomerifera, C. hypericifolia, E. glomerifera Chamaesyce melanadenia
Name authority Linnaeus: Sp. Pl. 1: 454. (1753) Torrey: in War Department [U.S.], Pacif. Railr. Rep. 4(5): 135. (1857)
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