Euphorbia hypericifolia |
Euphorbia geyeri |
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graceful sandmat, graceful spurge |
Geyer's sandmat, Geyer's spurge |
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Habit | Herbs, annual, with taproot. | Herbs, annual, with taproot. | ||||
Stems | erect to ascending, 15–50 cm, glabrous. |
prostrate or slightly ascending, loosely mat-forming, 4–25(–45) cm, glabrous. |
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Leaves | opposite; stipules connate, deltate, usually entire, sometimes laciniate-fringed at tip, 1.5–2.2 mm, glabrous; petiole 1–3 mm, glabrous; blade obliquely oblong-oblanceolate, 10–35 × 7–15 mm, base asymmetric, oblique, margins serrate or serrulate, especially toward apex, apex broadly acute, surfaces glabrous; palmately veined at base, pinnate distally. |
opposite; stipules usually distinct, occasionally connate basally on lower side, usually deeply parted into 3 or more filiform segments, 0.7–1.5 mm, glabrous; petiole 1–2 mm, glabrous; blade oblong to oblong-obovate or oblong-elliptic, 4–12 × 2–6 mm, base slightly asymmetric, angled or rounded, with one side usually expanded into small, rounded auricle, margins entire, apex usually truncate, occasionally emarginate, abaxial surface pale grayish green, both surfaces glabrous; only midvein conspicuous or venation obscurely pinnate (larger leaves). |
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Involucre | obconic, 0.9–1.1 × 0.4–0.9 mm, glabrous; glands 4, yellow-green to brown, stipitate, subcircular, 0.2 × 0.2 mm, occasionally nearly rudimentary; appendages absent on smaller glands or white to pink, shape highly variable, usually round to ± elliptic, 0.3–0.4 × 0.5–0.7 mm, distal margin entire. |
broadly campanulate, 1–1.5 × 0.7–0.9 mm, glabrous; glands 4, green to reddish, elliptic-oblong to nearly circular, slightly cupped to folded, 0.2–0.4 × 0.2–0.6 mm; appendages rudimentary to absent or white to reddish-tinged, usually rounded, sometimes pointed, (0–)0.5–1 × (0–)0.1–1.2 mm, distal margin entire or slightly toothed. |
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Staminate flowers | (0–)2–20. |
5–20. |
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Pistillate flowers | ovary glabrous; styles 0.4 mm, 2-fid 1/2 length. |
ovary glabrous; styles 0.2–0.6 mm, 2-fid nearly 1/2 length. |
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Capsules | depressed-globoid, 1.3–1.4 × 1.1–1.5 mm, glabrous; columella 1–1.1 mm. |
globose-ovoid, 1.5–2 × 1.5–3 mm, glabrous; columella 1.5–1.9 mm. |
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Seeds | with very thin whitish mucilaginous coat over light brown testa below, ovoid-triangular, bluntly 4-angled in cross section, 0.9–1.1 × 0.5 mm, with shallow irregular depressions alternating with low, smooth ridges. |
ashy white, ovoid, terete to bluntly subangled in cross section, 1.1–1.7 × 0.9–1.2 mm, smooth, with smooth brown line from top to bottom on adaxial side. |
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Cyathia | in dense, axillary and terminal, capitate glomerules with reduced, bractlike leaves subtending cyathia; peduncle 0.5–1.8 mm. |
solitary or in small, cymose clusters at distal nodes; peduncle 1–2 mm. |
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Euphorbia hypericifolia |
Euphorbia geyeri |
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Phenology | Flowering and fruiting early spring–late fall. | |||||
Habitat | Open, disturbed areas, nurseries. | |||||
Elevation | 0–200 m. (0–700 ft.) | |||||
Distribution |
AL; AR; FL; GA; LA; OK; SC; TX; Central America; South America; West Indies [Introduced in Asia, Pacific Islands]
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CO; IA; IL; IN; KS; LA; MI; MN; MO; MS; MT; ND; NE; NM; OK; SD; TX; UT; WI; WY; MB; n Mexico
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Discussion | Euphorbia hypericifolia is native to the New World tropics, and it is most likely adventive in the flora area (where it is most widely distributed in Florida and Texas). Reports from Arizona, California, and Maryland likely represent waifs or misidentifications. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). The two varieties of Euphorbia geyeri have been distinguished in large part by the presence of conspicuous involucral gland appendages in var. geyeri and the lack of appendages in var. wheeleriana. The two varieties are recognized here, but the variation in the size and presence of involucral gland appendages in the closely related E. bombensis suggests that this might be a somewhat variable character in this group of species. Euphorbia geyeri is widespread throughout the central United States in sandy soils. Populations at the eastern edge of the range are often considered adventive (for example, sandy soils along railroad grades in Michigan). Euphorbia geyeri resembles E. glyptosperma (both being entirely glabrous), but that species has serrulate leaves (near the apex) and strongly angled, transverse-ridged seeds whereas E. geyeri has entire leaves and smooth, rounded seeds. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 12, p. 272. | FNA vol. 12, p. 268. | ||||
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum | ||||
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Subordinate taxa | ||||||
Synonyms | Chamaesyce glomerifera, C. hypericifolia, E. glomerifera | Chamaesyce geyeri | ||||
Name authority | Linnaeus: Sp. Pl. 1: 454. (1753) | Engelmann: Boston J. Nat. Hist. 5: 260. (1845) | ||||
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