Euphorbia hypericifolia |
Euphorbia cumulicola |
|
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graceful sandmat, graceful spurge |
coastal dune sandmat |
|
Habit | Herbs, annual, with taproot. | Herbs, usually annual, rarely perennial, with taproot. |
Stems | erect to ascending, 15–50 cm, glabrous. |
prostrate, 10–20 cm, glabrous. |
Leaves | opposite; stipules connate, deltate, usually entire, sometimes laciniate-fringed at tip, 1.5–2.2 mm, glabrous; petiole 1–3 mm, glabrous; blade obliquely oblong-oblanceolate, 10–35 × 7–15 mm, base asymmetric, oblique, margins serrate or serrulate, especially toward apex, apex broadly acute, surfaces glabrous; palmately veined at base, pinnate distally. |
opposite; stipules distinct, linear-subulate, usually divided into 3–7 linear segments, 0.5–1 mm, glabrous; petiole 0.5–1 mm, glabrous; blade narrowly elliptic to lanceolate, 4–8 × 1–2 mm uniform in size, base asymmetric, obtuse to rounded, margins entire, apex obtuse to acute, surfaces green to reddish flushed, glabrous; only midvein conspicuous. |
Involucre | obconic, 0.9–1.1 × 0.4–0.9 mm, glabrous; glands 4, yellow-green to brown, stipitate, subcircular, 0.2 × 0.2 mm, occasionally nearly rudimentary; appendages absent on smaller glands or white to pink, shape highly variable, usually round to ± elliptic, 0.3–0.4 × 0.5–0.7 mm, distal margin entire. |
campanulate, 0.9–1.1 × 1–1.3 mm, glabrous; glands 4, green to red, slightly stipitate, subcircular, 0.1 × 0.1 mm; appendages white or pink, fringing edge of gland, short-flabellate, often rudimentary, 0.1–0.2 × 0.1 mm, distal margin crenate or entire. |
Staminate flowers | (0–)2–20. |
5–8. |
Pistillate flowers | ovary glabrous; styles 0.4 mm, 2-fid 1/2 length. |
ovary glabrous; styles 0.2–0.3 mm, 2-fid at apex to nearly 1/2 length. |
Capsules | depressed-globoid, 1.3–1.4 × 1.1–1.5 mm, glabrous; columella 1–1.1 mm. |
ovoid or subglobose, 1.8–2 × 2–2.3 mm, glabrous; columella 1.2–1.5 mm. |
Seeds | with very thin whitish mucilaginous coat over light brown testa below, ovoid-triangular, bluntly 4-angled in cross section, 0.9–1.1 × 0.5 mm, with shallow irregular depressions alternating with low, smooth ridges. |
white to gray-brown, ovoid, terete to bluntly subangled in cross section, 1–1.3 × 1 mm, smooth or minutely pitted, with smooth brown line from top to bottom on adaxial side. |
Cyathia | in dense, axillary and terminal, capitate glomerules with reduced, bractlike leaves subtending cyathia; peduncle 0.5–1.8 mm. |
solitary at distal nodes; peduncle 1 mm. |
Euphorbia hypericifolia |
Euphorbia cumulicola |
|
Phenology | Flowering and fruiting early spring–late fall. | Flowering and fruiting year-round. |
Habitat | Open, disturbed areas, nurseries. | Sandy oak hammocks, open sandy areas behind mangroves, disturbed sandy sites. |
Elevation | 0–200 m. (0–700 ft.) | 0–10 m. (0–0 ft.) |
Distribution |
AL; AR; FL; GA; LA; OK; SC; TX; Central America; South America; West Indies [Introduced in Asia, Pacific Islands]
|
FL |
Discussion | Euphorbia hypericifolia is native to the New World tropics, and it is most likely adventive in the flora area (where it is most widely distributed in Florida and Texas). Reports from Arizona, California, and Maryland likely represent waifs or misidentifications. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia cumulicola could be confused with E. bombensis, both of which are widespread in Florida, but there the latter occurs on beaches close to the ocean, whereas E. cumulicola is not a beach-inhabiting species but occurs in more protected sandy habitats such as hammocks or stabilized dunes behind mangroves. Also, E. bombensis has fleshier leaves and larger seeds, and is more compact in habit than E. cumulicola, which is a more sprawling and densely branched plant with leaves that are usually purplish tinged along the margins. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 272. | FNA vol. 12, p. 265. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Chamaesyce glomerifera, C. hypericifolia, E. glomerifera | Chamaesyce cumulicola |
Name authority | Linnaeus: Sp. Pl. 1: 454. (1753) | (Small) Oudejans: Phytologia 67: 45. (1989) |
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